Palhinhaea riofrioi (Sodiro) Holub (1991: 93)

Øllgaard, Benjamin & Testo, Weston, 2021, The Lycopodiaceae of Panamá, Phytotaxa 526 (1), pp. 1-66 : 58

publication ID	https://doi.org/ 10.11646/phytotaxa.526.1.1
DOI	https://doi.org/10.5281/zenodo.5784375
persistent identifier	https://treatment.plazi.org/id/286E8977-7B77-FD62-10A9-FC34CF5C4009
treatment provided by	Plazi
scientific name	Palhinhaea riofrioi (Sodiro) Holub (1991: 93)
status

Treatment

5. Palhinhaea riofrioi (Sodiro) Holub (1991: 93) View in CoL . Fig. 22B

Lycopodium riofrioi Sodiro (1893: 582) View in CoL .— Lycopodiella riofrioi (Sodiro) B.Øllgaard (1987: 176) View in CoL . Type:— ECUADOR: In silv. occid. m. Pichincha ad Gualea , 9/888, Sodiro s. n. ( SI 21230 holotype).

Lycopodium pensum Lellinger & Mickel ex Lellinger (1977: 717) View in CoL . Type:— COSTA RICA: Prov. Heredia, 6 mi. from San Rafael de Heredia on slopes of Volcán Barba [Cerro Chompipe], McAlpin 216 (DUKE holotype n.v.; GH isotype).

Plants with very short to long, arching-looping runner shoots, rooting at soil contact, these bearing dorsally arising, erect, to 2 m tall, tree-like branched shoots. Erect shoots with several subdecussate to alternate, 10–20 cm long, subequally branched, ultimately long pendulous lateral branchlet systems. Ultimate branchlets 2–3 mm in diam. incl. leaves. Branchlet leaves imbricate, densely crowded, borne in alternating whorls of 4–6, these 1–1.5 mm apart, forming 8–12 longitudinal ranks, lanceolate to ovate-lanceolate, acuminate, 2.5–4 × ca. 1 mm, with densely ciliatefimbriate margins, with a very short, patent, terete-subangular base. Stems and leaf bases densely hairy. Strobili sessile, terminating ultimate branchlets, 1–2.5 cm long, 3–5 mm in diam. incl. sporophylls. Sporophylls borne in alternating whorls of 5, forming 10 longitudinal ranks, with coalescent bases, almost enclosing the sporangia, with ca. 2.5 × ca. 1 mm, deltoid, short acuminate exterior face, with narrowly membranous, erose-laciniate margins. Sporangia 0.7–1 mm in diam.

Distribution: Costa Rica, Panamá, Venezuela and adjacent Brazil, Colombia, Ecuador, Peru.

Habitats: Wet montane forest, in clearings and on road banks, 1660–2240 m elev.

Specimens studied:— Chiriquí: Cerro Punta, Las Nubes PILA, sendero La Cascada , 2240 m, Florpan 6870 ( AAU, PMA). Cerro Colorado, along road N of San Felix, 18.5 mi N of bridge over river near San Felix, 6.5 mi beyond Chame at road to Escopeta , 1660 m, Croat 74985 ( MO, right hand specimen). Mining road 20 mi above bridge over Río San Felix , 2000 m, Antonio 2609 ( MO, PMA) . Chiriqui / Bocas del Toro: Trail along continental Divide , to ca. 3 km E of Cerro Pate Macho, 2000–2200 m, Smith et al. 2448 ( UC) .