Copelatus bacchusi Wewalka, 1981

Jiang, Zhuo-Yin, Zhao, Shuang, Yang, Zhen-Yu, Jia, Feng-Long & Hájek, Jiří, 2022, A review of Copelatus Erichson, 1832 of Mainland China, with description of ten new species from the japonicus complex (Coleoptera: Dytiscidae: Copelatinae), Zootaxa 5124 (3), pp. 251-295 : 253

Copelatus bacchusi Wewalka, 1981

( Figs 1 , 29–30 )

Copelatus bacchusi Wewalka, 1981: 66 .

Type locality. “Indien, Utar Pradesh, Kumaun, Haldwani” [ India, Uttarakhand State, Kumaon Division, Haldwani , ca. 29°13′N 79°31′E] .

Type material. Holotype ♂, deposited in BMNH (not studied).

Material studied. CHINA: Hainan: 1 ♂, Jianfeng Mts., Tian Chi , 750 m, 23.i.1996, Ji & Wang leg. ( NHMW) . Yunnan: 1 ♂, Xishuangbanna , ca. 15 km W Menglun, ca. 700-800 m, 5.xi.1999, M. Jäch et al. leg. ( NHMW) .

Diagnosis. Medium sized (TL: 4.9–5.3 mm), oblong-oval species. Head with testaceous clypeus, frons and vertex usually darkened, brown; pronotum orange-brown to dark brown, laterally broadly testaceous; elytra brown with testaceous lateral sides and broad basal testaceous band not reaching suture. Pronotum with short longitudinal strioles laterally. Each elytron with six complete discal striae and one submarginal stria: striae 1–5 beginning at base and ending close to apex, odd striae somewhat longer than even ones; stria 6 shorter, absent at base and ending at apical fourth; submarginal stria rather short, presenting only in third fourth of elytral length ( Fig. 1 ). Median lobe of aedeagus in lateral view weakly sickle-shaped with apex slightly skewed on ventral side ( Fig. 29 ). Parameres slender, ‘C’-shaped; apex short and broad; apical lobes long, club-shaped ( Fig. 30 ). Female similar to male, strioles on pronotum more numerous.

Distribution. The species was described, and for 40 years known only from the Uttarakhand State in northern India. Recently, Shaverdo et al. (2021) provided additional records not only from Bhutan and Nepal in the Himalayan Region, but also from Myanmar, Thailand and Vietnam, suggesting that C. bacchusi is widespread in the continental southeast Asia. The present records from two distant Chinese provinces confirm this statement. First records from China (Hainan and Yunnan provinces) ( Fig. 81 ).

Figures

FIGURE 81. Map of distribution of Copelatus in China Mainland.

FIGURES 1–4. Habitus of Copelatus in China. 1. C. bacchusi (♂; Xishuangbanna, Yunnan); 2. C. chinensis (♂; Lankeshan, Zhejiang); 3. C. chinensis (holotype ♀ of C. collocallosus); 4. C. dentatipenis sp. nov. (holotype ♂; Jinggangshan, Jiangxi). Scale bar 1 mm.

FIGURES 29–38. Male genitalia of Copelatus. 29, 31, 33, 35, 37. median lobe in lateral view; 30, 32, 34, 36, 38. paramere. 29–30. C. bacchusi (Hainan); 31. C. chinensis (a. Lakeshan, Zhejiang; b. Yanjiajie, Hunan; c. Tainmushan, Zhejiang); 32. C. chinensis (Tianmushan, Zhejiang); 33. C. dentatipenis sp. nov. (a. holotype, Jinggangshan, Jiangxi; b. Guadun, Fujian; c. Mangshan, Hunan); 34. C. dentatipenis sp. nov. (holotype, Jinggangshan, Jiangxi); 35–36. C. felicis sp. nov. (holotype, Baishui, Yunnan); 37. C. diversistriatus sp. nov. (a. holotype, Pu’er, Yunnan; aa. detail of lateral carina with spines; b. Attapeu, Laos; c. NE Mintaingbin Forest Camp, Myanmar); 38. C. diversistriatus sp. nov. (holotype, Pu’er, Yunnan). Scale bar 0.5 mm.