Quinquala obovata MANCHESTER et DISNEY, 2020

Manchester, Steven R., Disney, Kory A. & Pham, Kasey K., 2020, Winged Fruits Of Rutaceous Affinity From The Eocene Of Western North America, Fossil Imprint 76 (2), pp. 211-216 : 213

Quinquala obovata MANCHESTER et DISNEY sp. nov.

Text-figs 2 , 3a–g, i, j , 4

H o l o t y p e. UF 19376-60038 ( Text-fig. 2a ) housed in

Florida Museum of Natural History, Gainesville, USA.

P a r a t y p e s. UF 19376-60023a ( Text-fig. 2b ), UF 19374-60339 ( Text-fig. 2c ), UF 19376-60050 ( Text-fig. 2d ), UF 19374-61756 ( Text-fig. 2e ), UF 19376-60067 (Textfig. 2f, p), UF 19376-60069 ( Text-figs 2i , 3e ), UF 19376- 60072 ( Text-fig. 2j ), UF 19376-60062 ( Text-figs 2l , 3f ), UF 19374-60403 ( Text-fig. 2m ), UF 19376-60023b (Textfig. 3a), UF 19376-60023c ( Text-fig. 3b ), UF 262-17690 ( Text-fig. 3c, g ), UF 229-53091 ( Text-fig. 3d ) housed in Florida Museum of Natural History, Gainesville, USA.

P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.

PFN001528 (for new species).

E t y m o l o g y. The epithet, obovata refers to the fruit shape.

T y p e l o c a l i t y. Kisinger Lakes, northwestern

Wyoming, USA ( UF 19376 : N 43° 42.056′, W 109° 52.918′) .

T y p e h o r i z o n a n d a g e. Tepee Trail Formation,

Eocene.

A d d i t i o n a l l o c a l i t i e s. Kisinger Lakes ( UF 19374 : N 43° 42′ 01.9″, W 109° 52′ 44.9″; UF 19375 : N 43° 42′ 03.0″, W 109° 52′ 53.3″), West Branch Creek, northcentral Oregon, USA ( UF 229 : N 44° 34′ 53.40″, W 120° 15′ 57.31″; UF 230 : N 44° 35′ 25.31″, W 120° 15′ 27.83″; Eocene Clarno Formation), White Cliffs, northcentral Oregon ( UF 262 : N 44° 44.302′, W 120° 28.376′; Eocene Clarno Formation) .

D i a g n o s i s. Fruits single, obovate, 1.1–1.7 times longer than wide. Fruit apex rounded, without a stylar protrusion. Base cuneate and rounded, margins entire. Locular area oblanceolate. Five thick longitudinal wings, veins obscure. Wing and fruit body dotted with circular glands. Fruit borne on pedicel with prominent perianth scar at the junction of pedicel and fruit base. Narrow disk scar located immediately below the perianth scar.

D e s c r i p t i o n. Fruits single, obovate, 9–15 mm long and 6–11 mm wide with a length/width ratio of 1.1–1.7, avg. 1.4. Fruit apex rounded, without a stylar protrusion. Base cuneate and rounded, margins entire. Locular area oblanceolate. Five thick longitudinal wings, veins obscure. Wing and fruit body dotted with circular glands 80–110 μm, avg. 100 μm diameter. Fruit borne on pedicel 10–13 mm long and 0.6–1.3 mm thick, with prominent perianth scar at the junction of pedicel and fruit base. Narrow disk scar located immediately below the perianth scar.

Figures

Text-fig. 2. Quinquala obovata gen. et sp. nov., fruits from Kisinger Lakes flora, Wyoming. a: Lateral view of fruit on long pedicel. Note the two lateral wings and pair of longitudinal grooves representing wings extending into the matrix. UF 19376-60038. b: Lateral view of fruit on long, bent pedicel. Note the perianth scar at the base of the fruit (arrow). UF 19376-60023a. c: Lateral view of fruit on long, slightly bent pedicel. Note the pair of longitudinal grooves representing wings extending into the matrix, and

Text-fig. 3. a–g, i, j: Quinquala obovata gen. et sp. nov., fruits from Wyoming and Oregon. a, b: Kisinger Lakes flora, Wyoming. a: Enlarged lateral view of split wing displaying venation along outer edge. Note the seed present in the locular area (arrow). UF 19376-60023b. b: Enlarged lateral view of split wing containing seeds within the locular area (arrow). UF 19376-60023c. c: Lateral view of fruit from White Cliffs, Oregon. UF 262-17690. d: Lateral view of fruit from West Branch Creek, Oregon. UF 229-53091.

Text-fig. 4. Reconstruction of Quinquala obovata fruits; artwork by K. K. Pham.