Xestopus Andrewes, 1937,

Zhu, Pingzhou, Kavanaugh, David H. & Liang, Hongbin, 2021, Notes on the genus Xestopus from China, with description of a new species (Carabidae, Sphodrini, Dolichina), ZooKeys 1009, pp. 139-151: 139

publication ID

http://dx.doi.org/10.3897/zookeys.1009.61515

publication LSID

lsid:zoobank.org:pub:3E78899F-3A6F-4C21-A690-4375A15EA34A

persistent identifier

http://treatment.plazi.org/id/2AACC6DF-9C28-5942-89C2-5E4E529A077F

treatment provided by

ZooKeys by Pensoft

scientific name

Xestopus Andrewes, 1937
status

 

Genus Xestopus Andrewes, 1937 

Xestopus  Andrewes 1937: 59; Morvan 1979: 41 (mentioned only); Casale 1981: 389 (key to species); Morvan 1982: 45 (described new species); Casale 1988: 138 (placed in Dolichina  ); Sciaky and Facchini 1997: 235 (described new species); Sciaky and Wrase 1998: 223 (key to genera of Dolichina  ); Lorenz 1998: 373 (catalogue); Hovorka and Sciaky 2003: 530 (catalogue); Lorenz 2005: 399 (catalogue); Hovorka 2017: 769 (catalogue); Schmidt and Will 2020: 336 (diagnosis).

Nepalocalathus  Synonym: Nepalocalathus  Habu, 1973: 100, type species Calathus kumatai  Habu, 1973; Habu 1978: 302 (raised to genus rank and transferred to Dolichina  ); Casale 1981: 389 (synonymized with Xestopus  ).

Xestopus  Synonym: Wittmerosphodrus  Morvan, 1978: 100, type species Wittmerosphodrus walteri  Morvan, 1978; Morvan 1979: 36 (described a new species); Casale 1981: 389 (synonymized with Xestopus  ).

Type species.

Pristonychus alticola  Fairmaire, 1889 (type locality: Mountain Yeomatong, North Myanmar; holotype in NZSI), by monotypy.

Diagnosis.

Among the seven genera of Dolichina  , Xestopus  can be distinguished from others by the following character combination: third antennomere very long, usually longer than the first two antennomeres combined; tooth of mentum bifid; pronotum more or less cordiform; elytra with microsculpture nearly isodiametric, parascutellar seta present, elytral disc without setigerous pores; lateral grooves absent on metatarsomeres I-IV; males with right paramere hooked at apex; female apical gonocoxite with one ensiform seta at external margin (in most specimens) and sensory pit absent. Detailed descriptions and a key to the genera in the subtribe Dolichina  have been provided by Casale (1981) and Sciaky and Wrase (1998).

Comparisons.

This genus is most similar to the genus Dolichus  Bonelli, sharing the large body size (>15 mm), but its members differ from those of the latter in having the tooth of the mentum bifid, the pronotum cordiform, elytral interval 3 without setigerous pores, and the right paramere of male genitalia apically hooked.

Distribution.

This genus includes eight species distributed along the Himalayas (two in Nepal, three in Bhutan, two in China, and one in Myanmar).

Remarks.

The type species of this genus, Xestopus alticola  (Fairmaire, 1889) was described from Mount Yeomatong, northern Myanmar. In the two versions of the Catalogue of Palearctic Coleoptera  ( Hovorka and Sciaky 2003; Hovorka 2017), the distribution of this species includes Sichuan and Sikkim and both these two are doubtful. The record for Sikkim was added by Andrewes (1937); however, this locality and the type locality are extremely distant from each other for a species with apterous members. In addition, the record for Sikkim maybe a different species ( Morvan 1979; Casale 1981). No other literature records have reported X. alticola  from Sichuan, and, in fact, no Xestopus  specimens have been found in that province during our many expeditions in Sichuan, so we also doubt the reliability of this distributional record.

Casale (1981) pointed out that the three species from Bhutan ( X. walteri  (Morvan, 1978), X. bhutanensis  (Morvan, 1979), and X. cordicollis  (Morvan, 1979)) may represent three subspecies of a single species or eventually be combined into one species when abundant materials are available. Conversely, the male allotype and female holotype of X. nepalensis  probably represent two different species, in our opinion, based on the original description and figures. The male differs from the female in having elytra with (1) rufous color in the scutellar region and along the sutural margin to mid-length, (2) the humeri more rounded, (3) the basal margination markedly sinuate, and (4) the apices more rounded and slightly oblique medially. Thus, a revision appears necessary to deal with this situation.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Carabidae

Loc

Xestopus Andrewes, 1937

Zhu, Pingzhou, Kavanaugh, David H. & Liang, Hongbin 2021
2021
Loc

Wittmerosphodrus

Morvan 1978
1978
Loc

Wittmerosphodrus walteri

Morvan 1978
1978
Loc

Nepalocalathus

Habu 1973
1973
Loc

Nepalocalathus

Habu 1973
1973
Loc

Calathus kumatai

Habu 1973
1973
Loc

Xestopus

Andrewes 1937
1937
Loc

Dolichina

Andrewes 1937
1937
Loc

Dolichina

Andrewes 1937
1937
Loc

Xestopus

Andrewes 1937
1937
Loc

Xestopus

Andrewes 1937
1937
Loc

Xestopus

Andrewes 1937
1937