Anisocentropus McLachlan, 1863

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 15

Anisocentropus McLachlan

Anisocentropus McLachlan, 1863: 492 .

Type species: Anisocentropus illustris McLachlan, 1863: 492 , subsequently selected by Ulmer (1929: 177).

Kizakia Iwata, 1927: 211 , 217; synonymized by Ulmer, 1951: 345)

Type species: Kizakia kawamurai Iwata, 1927: 211 , 217 (monobasic)

This genus is widely distributed, with many species described from the Australasian and Oriental biogeographical regions. It is also represented in the eastern part of the Afrotropical Region, including Madagascar, with few, less-known species, as well as a single species in the Nearctic Region and 1 Australasian-Oriental species ranging into the East Palearctic Region.

Malicky (1994) established 4 subgenera based primarily on the form of the preanal appendages:

(1) Anisocentropus s. str.: preanal appendages simple and oval.

(2) Anisokantropus: preanal appendages complex, subdivided.

(3) Anisolintropus : preanal appendages with supplementary small, filiform basal process.

(4) Anisomontropus: preanal appendages very long.

Additional subgeneric characters were presented in his subgeneric discriminatory matrix.

(1) The form of the forewing.

(2) The presence of a basal setal brush on the hind wing.

(3) The presence of a setal comb on each male hind leg.

(4) The tibial spur formula.

(5) The maxillary palp formula.

It appears that the 5 additional characters are systematically less important, secondary features that develop and become reduced sporadically and independently in unrelated species. We are of the opinion that the subgenera established by Malicky are simply diagnostic species-groups characterised by the genital structural elements, mostly the preanal appendages.

The gonocoxites are long and simple, each with a variously wide basal part in the following Oriental species: A. annulicornis ( Hagen, 1858) ; A. apis Malicky, 1998 ; A. atropurpurpurens Flint, 2000 ; A. bacchus Malicky & Chantaramongkol, 1994 ; A. cameloides Malicky, 1995 ; A. bungus , new species A. csorbai , new species A. diana Malicky & Chantaramongkol, 1994 ; A. flavomarginatus Ulmer, 1906 ; A. handsschini Ulmer, 1951 ; A. insularis Martynov, 1930 ; A. ittikulama Schmid, 1958 ; A. janus Malicky & Chantaramongkol, 1994 ; A. kawamurai (Iwata, 1994) ; A. kempi Martynov, 1936 ; A. longulus Navás, 1933 ; A. maculatus Ulmer, 1926 ; A. magnus (Banks, 1931) ; A. minutus ( Martynov, 1930) ; A. orion Mey, 1997 ; A. pyraloides (Walker, 1852) ; A. salsus (Betten, 1909) ; A. ulmeri Malicky, 1998 ; and A. thonmihn , new species.

In a few Oriental species the basal part of each gonocoxite is divided, with a produced dorsal lobe, as in: A. pandora Malicky & Chantaramongkol, 1994 ; and A. dvaupadhah , new species. The division of the gonocoxites in these 2 species is accompanied by simple preanal appendages.

One species with complex, divided preanal appendages, A. pan Malicky & Chantaramongkol, 1994 , has complex divided gonocoxites, each with 3 lobes.

Only 3 Oriental species have short gonocoxites: A. malaisei , new species A. erichthonios Malicky & Cheunbarn in Malicky et al., 2001; and A. golem Malicky, 1994 . Almost all known Anisocentropus species from the Australasian and Afrotropical Regions and from the Philippines (Oriental Region) have short gonocoxites. The single Nearctic species, A. pyraloides (Walker, 1852) is similar to the Oriental species in having long gonocoxites, with simple, elongated preanal appendages. Afrotropical species are rather poorly known, but the species all appear to have short gonocoxites. We examined recently collected specimens of A. voeltzkowi Ulmer, 1909 , and describe here a similar species, A. samuh , new species, both from Madagascar and having the same short gonocoxites, but each with a slender digitate process on its apicodorsal corner. A few species from the Australasian and Oriental Regions have the same short gonocoxites with slender digitate processes. Here we establish a new diagnostic species-group, Anisocentropus latifascia diagnostic species-group, based on the presence of short gonocoxites as well as presence of unusual lateral flanks on the posterior margins of segment IX. This diagnostic species-group includes the Anisocentropus brevipennis , new diagnostic speciescluster for species having short gonocoxites each with a slender, digitate process on its apicodorsal corner. The simple preanal appendages of the subgenus Anisocentropus (Anisocentropus) are modified in species from the Oriental region and include species with an elongated, robust structure in the subgenus Anisocentropus (Anisomontropus) and a complex structure in the subgenus Anisocentropus (Anisokantropus) , both subgenera established by Malicky (1994). Until more species are known, we place the A. latifascia diagnostic species-group into the subgenus Anisocentropus (Anisocentropus) Malicky, 1994 .