Anisocentropus (Anisocentropus) magnificus Ulmer, 1906

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 29-34

Anisocentropus (Anisocentropus) magnificus Ulmer

Figs 53–57

Anisocentropus magnificus Ulmer, 1906: 56 .

Type country: Philippines.

New Records: PHILIPPINES: Mindanao P. I., Mt. Apo School , 15 km SW Davao, 500 m, 22–31.x.1965 [D. Davis] — 2 males, 1 female ( NMNH) .

Anisocentropus (Anisocentropus) malaisei , new species

Figs 58–67

This species is medium-sized and brown with a slightly darker brown and broad subapical transverse band on the forewing. It belongs to a diagnostic species-group from the Philippines, Indonesia, and southern Australasia characterized by having short gonocoxites, and with representatives populating the Philippines, Indonesia, and islands further south into Australasia. A single species from Thailand, A. erichthonios Malicky & Cheunbarn also has short gonocoxites. Anisocentropus malaisei , new species is most similar to A. tapenan , new species, from the Solomon Islands. Anisocentropus malaisei is easily distinguished from that species by the presence of a lateral flank on the posterior margin of segment IX being small and rectangular, and not so large and triangular as in A. tapenan ; the preanal appendages are robust digitiform, not flattened; and the gonocoxites do not have the pronounced humps on the subapicomesal lobes.

Male (in alcohol). Body medium-sized, small-eyed, brown; legs, antennae and palps lighter; wings brown, each with broad, dark brown subapical transverse band. Head rectangular in dorsal view, almost as long as broad. Ocelli absent. Tentorium slender, without dorsal arms; posterior arms very short, robust, ending in pair of large posterior tentorial pits with strong tentorial bridge without anteromesal or posteromesal protuberances; anterior tentorial arms robust posteriorly and slender anteriorly, without median lamellate processes; broad posterior half ending in median keel in dorsal view and ventral corner in lateral view. Facial groove pattern modified, with plate-like flange or rim connecting anterior tentorial pits, theoretical lines separating frons and clypeus, frontogenal vertical groove, forming oblique, dorsal continuation from anterior tentorial pits, merging almost horizontally with broad antennal grooves; clypeogenal vertical grooves below anterior tentorial pits short, running slightly mesoventrad; conspicuous pattern of plate-like flanges and frontogenal and clypeogenal sutures forming frontogenal septum (complex of anterior tentorial arms). Small, inconspicuous subantennal grooves sinuous, running almost horizontally between large frontogenal compact warts and palpifers, forming proximal articulations of palpi on stipes; subocular grooves not visible; very short, pronounced frontal groove present between antennae, joining vertexal medioantennal compact setose warts. Vertexal groove pattern reduced, large anterad-directed, rounded elevation dominating anterior half of vertex, reaching interantennal area, delineated laterally by extremely large membranous antennal sockets with corrugated or granulous surface near frontogenal compact setose warts; stem of epicranial groove (coronal groove) vestigial; usually permanent antennal and ocular grooves accompanied by occipito-postgenal grooves, partly merging anterad, with or tangential to very large occipital compact setal warts. Labrum long, quadrangular, with rounded apex, freely hanging, sparsely setose. Mandibles long, slightly pigmented, oriented diagonally beside lateral edges of labrum, laciniae narrow, elongated, bearing few setae. Frontal setal warts absent on face, frontal interantennal warts absent due to forward directed vertexal elevation. Pair of large, nearly triangular frontogenal compact setal warts present on posterior pregenae, obliquely or almost horizontally from frontogenal grooves, alone representing warts visible on face. Forward directed, rounded triangular elevation on vertex bearing fused vertexal medioantennal compact setose warts anteriorly on head; vertexal lateroantennal compact setal warts absent; antennal socket enlarged. Pair of small, rounded vertexal ocellar compact setose warts and very small pair of vertexal medioocellar diffuse setose warts visible, with 1 or 2 setae in middle of vertex; obliquely located pair of large, ovoid occipital compact setose warts dominating posterior half of vertex; small pair of postgenal compact wart visible between posterior section of ocular grooves and large occipital compact setose warts. Maxillary palps damaged in holotype and paratype, filiform, maxillary palp formula incomplete: II-I-IV-III. Scapes rounded, long, about half as long as head; pedicels 2/5ths as long as scapes. Minute, rounded setose wart located proximally above articulation of each cervical sclerite on proepisternum. Large, compact setal warts present mainly on membranous part of cervix, touching anterior arms of cervical sclerites. Lateral cervical sclerites forming narrow anterior arms articulating anteriorly to back of head with occipital condyle above posterior tentorial pits and fusing with posterior cervical sclerites; posterior cervical sclerites forming narrow, elongate plates reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by thin ventral intercervical sclerites. Two pairs of pronotal warts present: large-sized dorsal pair transversely ovoid, elongate, with narrow ends almost touching mesally; second pair of small-sized, rounded setal warts located deep laterally, visible in lateral view. One pair of mesoscutal, diffused warts present, arranged in longitudinal lines along full length of mesoscutum, composed mostly of single, double and maximum triple setae in groups visible as weakly pigmented alveoli. Pair of mesoscutellar warts forming small, less-pigmented, rounded areas, each with 5 to 6 setal alveoli. Legs with symmetrical claws; spur formula 2, 4, 3; posteroapical spur on each foreleg twice as long as anteroapical spur; anterior spurs on midlegs 1/4th as long as posterior spurs; anteroapical spurs on hind legs 1/4th as long as posterior spurs. Forewings: length 8.0 mm, membrane brown, each forewing with darker brown, broad, subapical band, pale at middle; R1 running free along its length; base of discoidal cell located proximally of wing middle; forks I, II, III, IV, V present; crossveins h, sc -r, r, s, r -m, m, m -cu, and cu2 present; crossveins cu1 and cu -a absent; postanal vein absent. Hind wings: R1 meeting R2; forks I, II, III, V present; fork I longer than fork II.

Male genitalia. Abdominal segment IX fused annularly, without longitudinal groove separating dorsal and ventral parts; tergum as short as venter, dorsum and venter produced into narrow quadrangular processes in lateral view; anterior margins of segment IX convex, more developed in ventral half; posterior margins each forming large, regularly round plate with small additional flank below each preanal appendage. Antecosta weakly developed, forming narrow, marginal rim, equally thin along its entire length, without conspicuous external groove of antecostal suture; in dorsal view, tergum mesally convex, merging imperceptibly onto segment X; spine row absent on posterior margin of segment IX; entire segment smoothly glabrous, except larger subventral area. Intersegmental depression between segments IX and X gently sloping. Segment X longer than gonocoxites and preanal appendages, forming broad hood with ventrad-directed apical rim characterised by sinuous excavations visible in lateral, dorsal and ventral views, excavations forming 2 “teeth” pattern on ventrally curving margin; apicoventral setose lobes reduced to setose surfaces before apex; apicodorsal setose lobe with few tiny setae above middle of segment X. Dorsal interlobular gap narrow, deep, triangular. Preanal appendages digitiform in lateral and dorsal views; broadening slightly before apex, with weak constriction only basally. Gonocoxites without harpagones, triangular in lateral view, elongate and apically broadened in ventral view, apex abruptly narrowing. Phallic apparatus forming curved tube, dorsally convex, ventrally concave, sclerotized ventral apical lobe elongate, only part of phallicata visible above ventral lobe while retracted; phallotremal sclerite visible in lateral view, forming dorsad-curving sclerite in membranous phallicata; in ventral view, phallotremal sclerite complex, U-shaped with laterad-curving ends; slender ejaculatory duct distinctly reaching phallotremal sclerite complex.

Holotype male: MYANMAR: North East , Kambaiti, 6800 ft, 7.iv.1934, Malaise trap [R. Malaise] ─ ( NHML).

Paratype: Same data as holotype — 1 male ( NHML) .

Distribution: Myanmar.

Etymology: Malaisei, name given to tribute René Malaise, former entomologist at the Swedish Museum of Natural History, for his extremely productive collecting trip in Burma, and for constructing the Malaise trap.

Figures

FIGURES 53–57. Anisocentropus magnificus Ulmer, non-type male from the Philippines: Mindanao P.I. 53 — right forewing; 54 — genitalia, lateral; 55 — genitalia, dorsal; 56 — genitalia, ventral; 57 — phallus, lateral.

FIGURES 58–62. Anisocentropus malaisei, new species, holotype. 58 — head, dorsal; 59 — head, frontal; 60 — tentorium, dorsal; 61 — tentorium, lateral; 62 — right forewing.

FIGURES 63–67. Anisocentropus malaisei, new species, holotype. 63 — genitalia, lateral; 64 — genitalia, dorsal; 65 — genitalia, ventral; 66 — phallus, lateral; 67 — phalus, ventral.