Anisocentropus (Anisocentropus) fridae, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 20-23

Anisocentropus (Anisocentropus) fridae , new species

Figs 29–34

This medium-sized species is chestnut brown with yellow underside and legs, and is distinguishable from other species on the Fiji Islands by this contrasting chestnut and yellow color. The dark chestnut colored forewing membrane has a large and specially shaped median hyaline window. The median keel on tergum IX is the shortest among the related A. fijianus and A. hannahae , new species. The mesal margin of the gonocoxites is clearly convex in ventral view and different from the concave margins of the gonocoxites of A. fijianus and A. hannahae , new species.

Male (in alcohol). Body medium-sized, with chestnut brown forewing membrane; legs, antennae and ventral part of body yellow; palps, vertex, scuta and scutella chestnut brown; forewings each with indistinct pale patches around pterostigma and pale, traverse, narrow band at basal 1/3rd. Head rectangular in dorsal view, almost as long as broad. Ocelli absent. Tentorium not observed. Facial groove pattern modified by plate-like flange, or rim, connecting anterior tentorial pits, forming theoretical lines separating frons and clypeus; frontogenal vertical grooves forming dorsal continuation from anterior tentorial pits, oblique, almost horizontally merging with broad antennal grooves; clypeogenal vertical grooves located ventrally of anterior tentorial pits, short, ventrally directed mesad. Pattern of plate-like flange and frontogenal and clypeogenal sutures forming complex with anterior tentorial arms (frontogenal septum); small, poorly visible subantennal grooves running sinuously, almost horizontally between large frontogenal, compact warts and palpifers; subocular groove invisible; very short, pronounced frontal groove present between antennae and joining to vertexal medioantennal compact setose wart. Vertexal groove pattern reduced; large, anterad-directed, rounded elevation dominating anterior half of vertex, reaching interantennal area, delineated laterally by enlarged membranous antennal sockets; antennal sockets with corrugated or granulous surfaces near frontogenal compact setose wart. Stem of epicranial groove (coronal groove) vestigial; occipito-postgenal groove partly merging anteri- orlly, touching large, occipital, compact setal warts. Labrum vertically long, quadrangular, with narrowing, rounded apex, freely hanging, sparsely setose. Mandibles long, weakly pigmented, located laterally along labrum, lacinia narrow, elongate, bearing few setae. Frontal setal warts absent on face, frontal interantennal warts absent due to anterad-directed vertexal elevation. Pair of large, subtriangular frontogenal compact setal warts present on posterior pregenae, on obliquely or almost horizontal frontogenal groove, representing compact warts dominating face. Anterad-directed, rounded triangular elevation on vertex bearing fused vertexal medioantennal compact, anterodorsal setose wart. Vertexal, lateroantennal compact setal warts invisible due to enlarged antennal socket. Pair of small rounded vertexal ocullar compact setose warts and very small pair of vertexal medioocellar diffuse setose warts visible, with 1 or 2 setae in middle of vertex. Obliquely located, ovoid pair of large occipital compact setose warts dominating posterior half of vertex. Small pair of postgenal compact warts visible between posterior section of ocular grooves and large occipital compact setose warts. Maxillary palps broken on holotype and paratype; observed palp formula: II-I-III. Scapes rounded, long, 1/3rd as long as head; 2.5 times longer than pedicel. Two pairs of pronotal warts present; large-sized dorsal pair ovoid, transversely elongated, separated mesally by broad fissura; ovoid pair of small-sized, rounded, deeply laterally-located setal warts. One pair of mesoscutal diffused warts present, arranged in longitudinal lines running along entire mesoscutum, composed of 1, 2 or 3 setae on pale alveoli. Pair of mesoscutellar warts forming small, less pigmented, rounded areas, each with 5–6 setal alveoli. Minute, rounded setose warts present proximally on each side above articulation of cervical sclerite on proepisternum, even smaller than setal wart on precoxale. Large, compact setal wart present on each side mostly on membranous part of cervix, touching anterior arm of cervical sclerite. Lateral cervical sclerites forming narrow anterior arm articulating anteriorly with back of head, with occipital condyle above posterior tentorial pits and fused to posterior cervical sclerites; posterior cervical sclerite forming narrow elongated plate reaching prothoracic episternum, articulating with weakly sclerotized anteromedian band of prothoracic eusternum by slender ventral intercervical sclerites. Legs with symmetrical claws; spur formula 2, 4, 3; foreleg spurs equal; midleg anteroapical spur 1/6th as long as foreleg spurs, anterior subapical spur 1/3rd as long as foreleg spurs; hind leg anteroapical spur 1/3rd as long as foreleg spurs. Forewings: length 8.0 mm; membrane chestnut brown, nearly bare, with indistinct pale patches around pterostigma and pale traverse, narrow band at basal 1/3rd; large, hyaline, median, linear window present on vein M starting shortly before bifurcation of M, running to bifurcation of M1+2, continuing on crossveins r -m and s. R1 separate along its length; crossvein r connecting R1 and R2 near wing margin; discoidal cell base located at midpoint of wing; forks I, II, III, IV, V present; crossveins h, sc -r, r, s, r -m, m, m -cu, and cu2 present, crossveins cu1 and cu -a absent; postanal vein absent; nygma and thyridium darkly pigmented, large. Hind wings: R1 meeting R2; forks I, II, III, V present; fork I longer than fork II.

Male genitalia. Abdominal segment IX fused annularly, without longitudinal groove separating dorsal and ventral parts; tergum as short as venter, dorsum and venter narrowly produced in lateral view; anterolateral margin of segment IX triangular, slightly below middle of segment; posterior margin forming large regularly convex plate with well-developed lateral flank on each side between preanal appendage and gonocoxite; apical half of flank glabrous, less pigmented, slightly transparent. Antecosta weakly developed, forming narrow, marginal rim, equally thin along its entire margin, external groove of antecostal suture inconspicuous; tergum IX with slightly protruding mesal keel; short, obtusely angled triangle in dorsal view; spine row absent on posterolateral margin of segment IX, segment IX glabrous except for small dorsopleural and large ventropleural and ventral setose areas. Intersegmental depression between segments IX and X forming deep step in lateral view due to protruding mesal keel of tergum IX. Segment X much longer than gonocoxites, as long as preanal appendages and forming broad hood with ventrad- and laterad-directed apical rim characterised by sinuous excavations visible in dorsal and ventral view, excavations resulting in formation of tooth-like pattern on ventrad-curving margin, with 2 teeth; apicoventral setose lobes reduced to setose surface subapically; apicodorsal setose lobe forming feeble, sparsely setose surface above middle of segment X. Dorsal interlobular gap deep, triangular, narrowing anteriorly. Preanal appendages digitiform in lateral view; clavate in dorsal view, slender basally, broadened slightly subapically. Gonocoxites without harpagones, triangular in lateral view, each elongate with convex mesal margin in ventral view, apex slightly narrowed; 4 stout setae located on middle of mesal surface. Phallic apparatus forming curving tube, dorsally slightly convex, ventrally concave in lateral view, elongated, ventral apical lobe sclerotized, only part of phallicata visible above ventral lobe while retracted; phallotremal sclerite barely visible in lateral view as large compact structure in membranous phallicata, phallotremal sclerite complex in ventral view, clearly visible as V-shaped sclerite with ends curving slightly laterad; slender ejaculatory duct clearly visible, reaching phallotremal sclerite complex, sinuous in lateral view, straight in ventral view.

Holotype male: FIJI ISLANDS: Viti Levu : Naitasiri Province, Nakobalevu Mt., 22.ix–9.x.2002, 18 ° 03 ’ S, 178 ° 25 ’ E, rainforest, Malaise trap [M. Irwin, E. Schlinger & M. Tokota’a] — ( BMH, in alcohol).

Paratypes: same data as holotype ─ 1 female ( BMH, in alcohol); Naitasiri Province, 4.8 km N Veisari Stlmt., log road to Waivudawa, 12.xii.2002 – 3.i.2003, 18.075 ° S, 178.362 ° E, FBA 177102, Malaise trap [E. Schlinger & M. Tokota’a] ─ 2 males ( BMH, in alcohol); Pabitra Wabu Baseline Survey, 1034 m, 17– 20.xi.2003, Malaise samples collected from Delena Veikovi, - 17.5833°S, 178.0833°E ─ 2 females ( BMH, in alcohol); Naitasiri Province , 1.8 km E Navai Village , old trail to Mt. Tomaniivi (Victoria), 700 m, 6.vi– 15.vii.2003, Malaise trap: MO4 [E. Namatalau], - 17.621°S, 177.998°E ─ 1 female ( BMH, in alcohol); Vuda Prov. , Koroyanitu Pk., 1 km E Abaca Vlg., Savuione Trl. 800 m 17.667 ° S, 177.55 ° E, 20.xi–3.xii.2002, Malaise trap 1 [E. Schlinger & H. Tokota’a] – 2 females ( BMH, in alcohol); Vuda Prov. , Koroyanitu N. M. P., 0.5 km N Abaca Village , Malaise trap, 7–12.x.2002 [E. Schlinger & H. Tokota’a], 17.666°S, 177.550°E, 800 m – 1 female ( BMH, in alcohol); Vuda Prov. , 1 km E Abaca Vlg., Koroyanitu , Ntl. Pk. , 800 m, Savuione Trl., 17 ° 40 ’ S, 177.33 ° E, 26.x–5.xi.2002, Malaise trap [E. Schlinger & H. Tokota’a] – 1 female ( BMH, in alcohol) .

Distribution: Fiji Islands.

Etymology: fridae , named after Frida Svare Johanson, the name of the daughter of one of the authors (KAJ) of this species, name given to remember Frida’s 16 th birthday, representing the starting day of the work on this paper.

Figures

FIGURES 29–34. Anisocentropus fridae, new species, holotype. 29 — right forewing; 30 — genitalia, lateral; 31 — genitalia, dorsal; 32 — genitalia, ventral; 33 — phallus, lateral; 34 — phallus, ventral.