Lannapsyche suksma, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 109-111

Lannapsyche suksma , new species

Figs 245–248

This species is most close to L. chantaramongkolae , especially in having narrow harpagones. The reduced preanal appendages form a similarly concave setose surface on the border of segments IX and X. Lannapsyche suksma is distinguished from L. chantaramongkolae having larger body size; longer dorsum of segment IX; and non-tapering, elongate and bilobed segment X.

Male (in alcohol). Body medium; brown; legs, antennae and palps light brown; wings pale brown with paler setae. Head rectangular in dorsal view; about 2 times wider than long. Ocelli absent. Tentorium slender, without dorsal arm; posterior arms short and robust in dorsal view, ending in pair of large posterior tentorial pits; tentorial bridge strong, without anteromesal protuberance, with posteromesal hump; anterior tentorial arms slender, without median lamellate process, slightly broadening immediately before anterior apex. Facial groove pattern simple. Frontogenal vertical groove forming long dorsad continuation from anterior tentorial pits. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; short, running obliquely, almost horizontally laterad; subantennal grooves vertical, each with slightly mesad-turning anterior end, apparently not confluent to clypeogenal grooves. Vertexal grooves with thick rim of compact warts; stem of epicranial groove complete, dominating on vertex; occipito-postgenal groove anteriorly partly merging with skeletal ring of occipital setal warts. Labrum semi-circular, without setal warts; narrow anterior, weakly pigmented part movable, freely hanging. Frontal setal warts absent; frontal interantennal warts absent, or represented by rounded compact, elevated, posterior setal warts. Pair of rounded frontogenal, posterior, compact, setal warts present dorsally on pregenae, between vertical frontogenal and subantennal grooves. Pair of large, transverse, elongate, setose warts present below clypeogenal grooves, representing largest setal warts on facial area. Vertexal, lateroantennal compact wart rounded, located below antennal grooves; pair of vertexal medioantennal compact setose warts rounded, elevated on anterior ending of coronal groove, separated by deep cleft. Paired frontal grooves nearly invisible. Pair of large, occipital compact warts cover posterior half of head dorsum, almost fusing with vertexal ocellar compact setal warts. Vertically elongated postgenal compact wart curving along posterior section of ocular grooves. Maxillary palp formula I-V-IV-II-III; covered by long setae; each basal segment with subapicomesal nodule with setose apex; base of nodules somewhat membranous, indicating flexibility to eversion. Each scape slightly longer than head. Each pedicel shorter than first segment of each flagellum. Pronotum with 2 pairs setal warts. Mesal pronotal pair compact, slightly elongate obliquely, occupying top of dorsally elevated area; elevated area separated by wide gap. Lateral pronotal warts small, rounded. Pair of mesoscutal warts arranged in longitudinal lines composed of 4 to 5 fragments. Pair of mesoscutellar diffuse warts forming longitudinal patch. Each proepisternum with large, ovoid setose wart. Each precoxal wart small. Large, compact setal wart present on membranous part of cervix, tangential with narrow anterior arm of each cervical sclerite. Lateral cervical sclerites forming narrow anterior arms articulating anteriorly with occipital condyle above posterior tentorial pits on head; fusing with posterior cervical sclerites; each posterior cervical sclerite forming large, quadrangular plate reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerite. Leg claws symmetrical; spur formula 2, 4, 4; foreleg posteroapical half as long as anteroapical spur; midleg and hind leg posterior spurs 2 times longer than anterior spurs. Forewings: length 9.9 mm; membrane light brown without visible pattern, with pale setae; R1 confluent with R2 short before C; base of discoidal cell located proximally of mid-length of wing; forks I, II, III, V present; forks I, II, and V sessile, fork III petiolate, crossveins ic, r, s, r-m and m-cu present; crossvein m absent; crossveins arranged in regular vertical line of anastomis; postanal vein present. Hind wings: R1 confluent with R2 well before C.

Male genitalia. Segment IX fused annularly, longitudinal groove separating dorsal and ventral parts, continuing along ventral margin of segment X as dark rim; tergum concave, descending posteriad, about 2 times longer than sternum; anterior margins of segment IX slightly concave; posterior margins almost straight, with small triangular at end of longitudinal grooves. Antecostae forming weak marginal rims; thick ventrally of longitudinal grooves; external groove of antecostal sutures difficult to discern; tergum produced into central, sharp triangular in dorsal view, delineating concave surface of reduced preanal appendages; spine rows on posterior margins of segment IX reduced, merged with, or replaced by setae of preanal appendages. Intersegmental depression between segment IX and segment X visible, represented by rounded excision in lateral view. Segment X forming broad lobe in lateral view; with bilobed apex, dorsal lobe smaller in lateral view. Segment X with constricted base in dorsal view; apex bifid. Apicoventral setose lobes and apicodorsal setose lobes fused into integrated setose area on dorsum; composed of short, stout setae. Dorsal interlobular gap shallow, narrow. Superior appendages reduced into large circular, concave surface dominating on genitalia, covered by long setae. Each coxopodite longer than apex of segment X; parallel-sided, stout, straight. Harpagones slender, embedded into deep apical gap of coxopodites, apex covered with black stout and short triangular setae. Phallic apparatus forming inverted V; dorsal margin nearly straight in lateral view; almost as long as each coxopodite; sclerotized basal section long, curving ventrad, continuing into obscurely membranous part, possibly representing endotheca; more strongly sclerotized dorsally; ending in short, dorsal membranous phallicata being slightly narrower than preceding sclerotized part, and ventrally into ventral lobe being densely packed by microtrichia. Ejaculatory duct ending in circular phallotremal sclerites in endophallus. Strongly pigmented structure of phallic apparatus curving verticad in lateral view.

Holotype male: MYANMAR: North East : Kambaiti, 2000 m, 15.v.1934, Malaise trap (Malaise B.M. 1938-258) ─ ( NHML).

Distribution: Myanmar.

Etymology: Suksma, slender in Sanskrit, name referring to the slender harpagones.

Figures

FIGURES 245–248. Lannapsyche suksma, new species, holotype. 245 — genitalia, lateral; 246 — genitalia, dorsal; 247 — gonocoxite, ventral; 248 — phallus, lateral.