Lannapsyche birathena, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 107-109

Lannapsyche birathena , new species

Figs 241–244

This species resembles L. chantaramongkolae , from which it is easily separated by the presence of elevated setose surfaces at the border between segment IX and segment X, these structures are concave in L. chantaramongkolae . Furthermore, segment X is more deeply cleft in dorsal view, and the harpagones are broader in lateral and ventral view. The phallic apparatus is more complex in L. birathena than in L. chantaramongkolae .

Male (in alcohol). Body medium sized; brown; legs, antennae and palps light brown; wings pale brown with paler setae. Head rectangular in dorsal view, about 2 times broader than long. Ocelli absent. Tentorium slender, without dorsal arm; posterior arms short and robust in dorsal view, ending in pair of large posterior tentorial pits with strong tentorial bridge without anteromesal protuberance; posteromesal hump present; anterior tentorial arms posteriorly and anteriorly slender, without median lamellate process; slightly widening immediately before apex. Facial groove pattern simple. Frontogenal vertical grooves forming long, dorsal continuation of anterior tentorial pits. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; short, running obliquely, almost horizontally laterad. Subantennal grooves running vertically, anteriorly slightly turning mesad, not confluent to horizontal clypeogenal groove. Vertexal groove system simple, rim of compact warts thick; stem of epicranial grooves complete, dominating over entire vertexal area; other groove visible on vertex is only occipito-postgenal groove being partly merged anterad, with skeletal ring of occipital setal warts. Labrum semicircular-shaped, without setal warts; anterior, narrow, little pigmented part movable, freely hanging. Frontal setal warts absent. Frontal interantennal warts absent or possibly homolog with pair of rounded, compact setal warts located on elevated humps posteriorly on vertex. Pair of round frontogenal posterior compact setal warts present at dorsum of pregenae, between vertical frontogenal and subantennal grooves. Pair of large, transverse, elongate setose warts present below horizontal clypeogenal grooves; representing dominating setal warts on facial area. Lateroantennal vertexal compact warts rounded, located below each antennal groove. Pair of vertexal medioantennal compact setose warts rounded, visible at anterior end of coronal grooves, elevated, separated by deep fissure. Paired frontal grooves nearly invisible. Pair of large, occipital compact warts cover posterior half of head dorsum, almost fusing with vertexal ocellar compact setal warts. Vertically elongate, postgenal compact warts curving along posterior section of ocular grooves. Maxillary palp formula I-V-(II,III,IV); all segments covered by long setae; each basal segment with subapicomesal nodule with setose apex; base of nodules somewhat membranous, indicating flexibility to eversion. Each scapes slightly little longer than head. Each pedicel shorter than first segment of each flagellum. Pronotum with 2 pairs setal warts. Mesal pronotal pair compact, slightly obliquely elongate, occupying entire top of dorsad elevated area, elevated area separated by wide gap. Lateral pronotal warts small, rounded. Mesoscutum with pair of fragmented warts arranged in 2 longitudinal lines, each with 4 to 5 wart fragments. Each proepisternum with large, ovoid setose wart. Each precoxae with a small wart. Large, compact setal wart present on membranous part of cervix, tangential with narrow anterior arm of each cervical sclerite. Lateral cervical sclerites forming narrow anterior arms articulating anteriorly with occipital condyle above posterior tentorial pits; fused with posterior cervical sclerites; each posterior cervical sclerite forming large, quadrangular plate reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerite. Leg claws symmetrical; spur formula 2, 4, 4; posterior spurs on forelegs, and anterior spurs on midlegs and hind legs half as long as posterior spurs. Forewings: length 7.2 mm; membrane light brown, without visible pattern, with pale setae; R1 confluent with R2 shortly before C; base of discoidal cell located proximally of mid-length of wing; forks I, II, III and V present; forks I, II and V sessile, fork III petiolate; crossveins r, s, r-m and m-cu present; crossvein m absent; crossveins arranged in an almost regular line of anastomis; postanal vein present. Hind wings: R1 confluent with R2 well before C.

Male genitalia. Segment IX fused annularly; longitudinal groove separating dorsal and ventral parts, continuing along ventral margin of segment X as dark rim; tergum concave, slightly longer than sternum; anterior margins of segment IX ventrally concave; posterior margins triangular, with small, triangular at end of longitudinal groove. Antecostae weakly developed as marginal rims, thick ventrally of longitudinal groove; external grooves of antecostal suture unclear; tergum produced into central, triangular lobe in dorsal view; spine rows on posterior margins of segment IX reduced, merged with setae of preanal appendages. Intersegmental depression between segment IX and segment X well developed, stepwise. Segment X forming broad lobe, nearly parallel-sided, apex narrowing into small hump in lateral view. Segment X gradually narrowing from broad base in dorsal view; with bifid apex. Apicoventral setose lobes and apicodorsal setose lobes fused into integrated setose area on dorsum; composed of very short, stout setae. Dorsal interlobular gap triangular, deep, narrow. Superior appendages reduced into abbreviated, convex elevation covered by long setae. Each coxopodite longer than apex of segment X; parallel-sided, stout, straight. Harpagones broad, embedded into deep apical gap of coxopodites, widening apex covered by stout, black and short, triangular setae. Phallic apparatus as long as coxopodites; basal part long, curving ventrad, continuing into membranous part, possibly representing endotheca, followed by slightly sclerotized tube and membranous phallicata; ejaculatory duct conspicuous, running into circular, heavily striped endophallus. Phallotremal sclerites large, elongated, slightly bifid in lateral view.

Holotype male: MYANMAR: Kambaiti : 1800 m, 23.vi.1934, Malaise trap (Malaise B.M. 1938-258) ─ ( NHML).

Distribution: Myanmar.

Etymology: Birathena, broad in Sanskrit, name referring to the broad harpagones of this species.

Figures

FIGURES 241–244. Lannapsyche birathena, new species, holotype. 241 — genitalia, lateral; 242 — genitalia, dorsal; 243 — gonocoxite, ventral; 244 — phallus, lateral.