Psilotreta malickyi, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 102-105

Psilotreta malickyi , new species

Figs 232–237

This species closely resembles P. trimeresuri (Malicky) from Thailand. Psilotreta malickyi is separated from P. trimeresuri by having shorter preanal appendages, i.e. not longer than the end of the central body of segment X; by its dorsal interlobular gap being deep and widely triangular, not U-shaped; by the glabrous lateral plate that is long triangular in lateral view, reaching apex of the median bifid process of segment X; and by the presence of a basally broad, deltoid processes at the lateral face of segment X.

Male (in alcohol). Body large; light brown; legs, antennae and palps lighter than body; forewings pale brown. Head rectangular in dorsal view, about 2 times broader than long. Ocelli absent. Tentorium slender, with short, tapering dorsal arm; posterior arms short, thick in dorsal view, ending in pair of large posterior tentorial pits; with thick tentorial bridge with small anteromesal protuberance, without posteromesal hump; anterior tentorial arms slender, with small median lamellate process; slightly broadening before anterior apices. Facial groove pattern with deep indentation dominating on frons, continuing and broadening on vertex. Frontogenal vertical groove very short, almost reduced. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; short, running obliquely laterad. Subantennal grooves running vertically, close to ocellar grooves, slightly turning mesad; each with anterior part apparently confluent to oblique, almost horizontal clypeogenal groove. Subocular grooves merging to anterior meeting point of subantennal and clypeogenal grooves. Vertexal groove system with large indentation almost dominating on vertex, surrounded by frontal groove on face and by large, elongated lateral vertexal groove on vertex. Depression containing 4 rows of elongated, scaloid setae on bottom; stem of epicranial groove vestigial; visible posteriorly in depression; occipito-postgenal grooves partly merging anterad; tangential with lateral vertexal groove. Labrum small, rounded, without setal warts; long, narrow anterior part movable, freely hanging. Mandibles small, weakly pigmented, laciniae narrowly elongate. Frontal setal and frontal interantennal warts absent. Pair of narrow, elongate frontogenal compact setal warts present from antennal grooves, orienting anterad along subantennal groove of pregenae, representing only compact warts visible on face. Deep indentation nearly occupying entire vertex; vertexal lateroantennal compact setal warts slightly elongate obliquely, touching antennal grooves. Vertically elongate postgenal compact wart curving along posterior section of ocular grooves. Maxillary palp formula I-II-III-V-IV, each basal segment with small, well-defined setose subapicomesal nodule; each second segment very robust, covered by long setae, especially on apicodorsal half. Each scapes slightly little longer than head. Pedicels shorter than first segment of flagellae. Pronotum with 1 pair transversely elongate, broad warts dominating on entire dorsum. One pair mesoscutal, rounded, compact setose warts present. Mesoscutellar warts absent. Each proepisternum with large, ovoid, setose wart. Precoxal warts small. Large, compact setal wart present on membranous part of cervix, touching anterior arm of cervical sclerites. Lateral cervical sclerites forming narrow anterior arms articulating anteriorly with back of head at occipital condyle above posterior tentorial pits; fusing with posterior cervical sclerites; posterior cervical sclerite forming large, quadrangular plate reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerites. Leg claws symmetrical; spur formula 2, 4, 4, foreleg posteroapical spur shorter than anteroapical spur; midleg anterior spurs smaller than posterior spurs; hind leg spurs equally long. Forewings: length 14.5 mm; membrane light brown without visible pattern; R1 running separately to C, not confluent with, or recurrent into R2; base of discoidal cell located proximally of midlength of wing, originating near wing base; fork I almost twice as long as fork II; forks III and IV absent; forks I and II petiolate; crossveins r,s, r-m and cu-a present; crossveins m and cu absent; M reduced to single longitudinal vein with base drawn to R4+5 by short crossvein r-m, merging to base of fork V; postanal vein present. Hind wings: R1 running separate to C, fork I about 2 times longer than fork II.

Male genitalia. Abdominal segment IX fused annularly, with 2 incomplete longitudinal grooves separating dorsal, mesal and ventral parts; upper grooves straight; lower grooves slightly oblique dorsally; both grooves distrofied posteriorly on both sides; tergum straight, flat, with dorsad oblique rim in lateral view; half as long as convex venter in lateral view. Anterior margins of segment IX produced into large, subquadrangular lobe in; posterior margins straight vertical below segment X, with small flat lobe on ventral half. Antecostae weakly developed, each forming slender marginal rim, thicker on anterior lobes, without external grooves of antecostal sutures. Tergum with lateral, transverse rims continuing anteriorly, forming triangular in dorsal view; spine rows absent on posterior margins of segment IX; segment smoothly glabrous. Intersegmental depression between segment IX and segment X filled by basis of lateral plates of segment X. Segment X long, triangular, with long, median, dorsal process and setose, bifid apex; lateral plates glabrous; segment X constricted at base, continuing to long, narrow median process. Lateral plates deltoid. Apicoventral setose lobes present at apex of bifid median body. Apicodorsal setose lobes forming pair of rounded setose warts dorsally at very base of segment X, easily seen in dorsal view. Dorsal interlobular gap wide, deeply triangular. Superior appendages long, filiform in lateral view; digitiform in dorsal view. Coxopodites slightly shorter than apex of segment X; produced into long ventral and short dorsal branch. Harpagones small, originating mesally on point of bifurcation of each coxopodite; surface glabrous, with short, stout apical setae. Phallic apparatus large, straight; basal part slightly curving ventrad. Phallotheca produced ventrad. Endotheca short, with 2 pairs long spines visible inside phallotheca; apical half of spines curving dorsad. Small, curving phallotremal sclerite visible at proximal part of phallicata.

Holotype male: MYANMAR: North East , Kambaiti, 2000 m, 12.v.1934, Malaise trap (Malaise B.M. 1938-258) ─ ( NHML) . Paratype: same data as holotype ─ 1 female. Distribution: Myanmar. Etymology: Malickyi, named after Dr. Hans Malicky for his great contribution by describing the first species in this diagnostic species-group .

Figures

FIGURES 232–237. Psilotreta malickyi, new species, holotype. 232 —head, frontal; 233 —head, dorsal; 234 — genitalia, lateral; 235 — genitalia, dorsal; 236 — gonocoxite, ventral; 237 — phallus, lateral.