Phraepsyche yitungshana, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 117-120

Phraepsyche yitungshana , new species

Figs 264–271

This species is easily distinguished from the other 2 species in the genus in the forewings, particularly by the presence of a diffuse-spotted marginal and submarginal area; and in the genitalia by the intersegmental depression between segments IX and X without steps in lateral view. The phallic apparatus has a subapical, bulbous broadening being absent in the other 2 species. The apicoventral setose lobes on segment X are not widening laterally before apex in dorsal view, as in P. danaos ; some widening present at mid-length. The pronotum has only 1 pair of setal warts, and there are significant differences between the species also in the groove and setal wart patterns on the head.

Male (in alcohol). Body medium sized; light brown or yellowish; forewings light brown, with small spots on marginal and submarginal area. Head rectangular, almost 2 times broader than long. Ocelli absent. Tentorium without dorsal arm; posterior arms short, robust in dorsal view, ending in pair of large posterior tentorial pits, with strong tentorial bridge armed with small, 2-rayed anteromesal protuberance; anterior tentorial arms slender posteriorly, broadening anteriorly, with additional median lamellate process. Facial groove pattern complex. Frontogenal vertical groove long, with additional small arm running obliquely dorsomesad. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; short, running obliquely laterad. Subantennal grooves weak, running obliquely mesad, confluent to vertical frontogenal groove. Additional lateral grooves absent. Vertexal groove system with thick rim of compact warts; stem of epicranial grooves clearly visible on posterior margin of vertex. Paired groove running along middle of anterior part of vertex; most pronounced pair of grooves on vertex are sigmoid lateral vertexal grooves, nearly tangential in middle. Occipito-postgenal groove partly merging with skeletal ring of occipital setal warts, running along and reaching ocular grooves.

Labrum almost double-circuloid, with anterior, narrow, less pigmented part movable, freely hanging. Broad posterior, basal, sclerite of labrum bearing transversely elongate, compact setal wart. Pair of large frontal, lateral compact setal warts present on posterior end of frons; small pair of frontal, lateral, fragmented setose wars present on lower anterior end, comprising of 2 to 3 small warts; frontal interantennal warts absent, or possibly represented as larger anterior pair of warts. Pair of rounded frontogenal, posterior, compact setal warts present on dorsum of pregenae, tangential with subantennal grooves. Pair of frontogenal, lower, anterior, diffused setose warts present, with 4 to 5 individual setae. Vertexal, lateroantennal, compact setal warts elongated longitudinally, located mesally of lateral vertexal groove. Vertexal medioantennal diffuse setose warts absent. Pair of large occipital compact setal warts cover posterior half of head dorsum, almost integrating vertexal ocellar compact setal warts. Vertically elongated postgenal compact wart curving along posterior section of ocular grooves. Maxillary palps digitiform; maxillary palp formula I-II-(III,IV)-V, each with first segment more robust and broader than other segments, without subapicomesal nodule. Each scape almost as long as head. Each pedicel slightly longer than first segment of each flagell; basilateral part of first 13 segments of flagellae bearing long, branch-like processes. One pair pronotal warts present; lateral pronotal warts possibly fused with mesal warts; warts narrow, transversely elongate, occupying almost entire dorsum. Pair of mesoscutal compact, small, longitudinally elliptical warts present. Pair of longitudinally elongate, mesoscutellar compact warts present. Each proepisternum with large, ovoid setose wart. Precoxal warts small. Small, elevated protuberance-like compact, setal wart present on each cervical sclerite. Claws symmetrical. Spur formula 1, 4, 4; midleg anterior spurs half as long as posterior spurs; hind leg spurs almost equally long. Forewings: broad, 5.0 mm; membrane light brown, with diffuse pattern of small spots on costal, subcostal, radial, sub radial and first apical cell; Sc and R1 meeting in callosity as both veins thickening and fusing into broad, diffusion area at pterostigma; slightly elevated and translucent; forks III and IV absent; fork I and fork II long; meeting at pentagonal discoidal cell; M reduced; fork V wide; Cu2 absent; postanal vein present. Hind wings: Sc and R1 confluent at middle of wing; RS reduced, with stem running into M. Abdomen with sternal projection present on posterior margins of segment VI; forming elongated, robust rod (hammer) reaching segment margin. Hammer reinforced by ventrad curving internal ridge of longitudinal sutures.

Male genitalia. Abdominal segment IX fused annularly, short; tergum as long as sternum; anterior margins triangular, with sharply angled process located sub centrally; posterior margins concave. Antecostae well developed on sharply angled process; external groove of antecostal sutures well-developed; tergum without central triangular lobe in dorsal view; spine rows on posterior margins of segment IX highly reduced, with 2 to 3 setae near gonocoxites. Intersegmental depression between segment IX and segment X absent; tergum of segment IX continuing into segment X, without visible intersegmental step in lateral view. Segment X long in lateral and dorsal views, nearly parallel-sided; slightly triangular in lateral view; narrowing apically. Segment X slightly broadening from midway in dorsal view. Apicoventral setose lobes with several stout apical spines. Apicodorsal setose lobes not reaching middle of segment, fused into medial hump. Dorsal interlobular gap between apicoventral setose lobes deep, narrow. Superior appendages as broad in dorsal view as in lateral view; almost reaching medial hump of apicodorsal setose lobe. Coxopodites longer than apex of segment X, forming long, broad rods, slightly sigmoid in lateral view; curving mesad in ventral view. Harpagones slender, claw-like, both arising mesally, well before apex of coxopodites. Phallic apparatus shorter than each coxopodite, basal section sclerotized, short, curving ventrad, with short flange articulating with base of gonocoxites. Tubular phallotheca straight, with slightly convex dorsum, continuous with nearly membranous, spinefree phallicata. Weakly pigmented continuation of phallotheca obscure. Endotheca indistinct, straight, apex triangular. Gonopore indistinct, ejaculatory duct conspicuous, ending in circular and well pigmented anterior part of phallotremal sclerite. Dorsum of apex excavated, concave; posterior, narrow elongation of circular phallotremal sclerites visible, exposed in apicodorsal cavity.

Holotype male: CHINA: HONG KONG: Lantau South Country Park , stream 1.3 km E Yi Tung Shan Mt., Malaise trap, loc. 3, 20–27.v.2005 [A. Olsson & E. Ohlsson] ─ ( NRM).

Paratypes: same data as holotype ─ 1 male, 1 female ( NRM) .

Distribution: Hong Kong.

Etymology: Yitungshana, name derived from the local name of the type locality.

Figures

FIGURES 264–266. Phraepsyche yitungshana, new species, holotype. 264 —head, frontal; 265 —head, dorsal; 266 — head, lateral.

FIGURES 267–271. Phraepsyche yitungshana, new species, holotype. 267 — right wings; 268 — genitalia, lateral; 269 — genitalia, dorsal; 270 — gonocoxite, ventral; 271 — phallus, lateral.