Marilia jonssoni, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 78-81

Marilia jonssoni , new species

Figs 178–184

This species has a dark chestnut brown body colour, and together with the characteristic shape of the phallic apparatus, it resembles M. malickyi , new species. Marilia jonssoni is easily separated from M. malickyi , and other species, by the presence of highly enlarged first and second axillary sclerites on all wings, as well as having narrower vertex and small, almost rudimentary, medioantennal and lateroantennal compact setose warts.

Male (in alcohol). Body medium-sized; chestnut brown; legs, antennae and palps light brown; cephalic and thoracic setal warts white; wing membrane uniformly light grey with scattered dark gray setae. Eyes large, black; vertex constricted; distance between eyes about 1/5th the eye width; inter-ocular area 5 times longer than broad. Ocelli absent. Tentorium unknown. Facial groove pattern reduced; with pair of short, wide frontal lateral compact setose warts located strongly anteriorly; with fragmented white alveoli or setal thecae. Antennae with small antennal sockets on frons. Anterior remnant of frontal grooves invisible. Frontogenal vertical groove between frontal lateral compact warts and frontogenal compact setose warts invisible on untreated head. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; running obliquely laterad, reaching subgenal process. Subantennal grooves running vertically in narrow strip between ocellar groove and frontogenal compact setose warts; subocular groove not discernible. Vertexal groove pattern simplified, constricted; coronal groove almost complete, weakly discernible. Postoccipital groove encircling foramen magnum, or occipital foramen and postoccipital lobes. Labrum small, short, without setal warts, with movable anterior part, freely hanging. Mandibles almost indiscernible on uncleared head; lacinia visible, broader, setose. Pair of vertexal lateroantennal compact setal warts on constricted vertex small, round, elongate, dropshaped, posterior end narrower, following shape of compressed vertex. Vertexal mediantennal compact setose warts nearly tangential. Occipital compact setose warts diminished, nearly triangular at posterior end of coronal groove, following ocular groove of compressing eyes. Pair of sensilla basiconicae located at end of coronial grooves, almost tangential with occipital warts. Pair of postgenal compact warts following postgenal areas, curving along posterior part of ocular grooves, narrow strips pressed to ocular grooves. Postgenal surfaces glabrous, hidden by eyes. Maxillary palps slightly darker than legs and antennae; maxillary palp formula V- (II, III, IV)-I; each basal segment without setose subapicomesal nodule. Antennal scapes shorter than head. Pedicels shorter than first segment of flagellae. Pair of pronotal warts present, with white surfaces, covered by setae; transversely elongate, occupying entire pronotum, almost touching mesally, narrowest laterally. Second pair of pronotal warts small, ovoid, well separated from mesal warts. White mesoscutal diffuse setose warts arranged in longitudinally elongate, short, narrow, irregular patches on middle of dorsal surface of mesoscutum; separated by median notal groove. Median notal groove white, widening at mid-length. Narrow pair of lateral and oblique white bands continuing on apical half of mesoscutum. Almost entire surface of mesoscutellar areas white, covered by diffuse, white setal warts separated by median notal groove. Each proepisternum with ovoid setose wart being smaller than setose wart on each precoxale. Large, compact, setal wart present anteriorly on cervical sclerites; apparently forming sclerotized surface on membranous part of neck, touching only anterior cervical sclerite. Lateral cervical sclerites forming narrow anterior arm articulating anteriorly to back of head with occipital condyle above posterior tentorial pits, fused to posterior cervical sclerites. Posterior cervical sclerite forming posterad broadening plate reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerites; dark pattern of cervical sclerite complex clearly visible on pale membranous neck. Legs claws symmetrical; spur formula 2, 4, 4; forelegs with anteroapical spur half as long as posteroapical spur. Midlegs with apical posterior spur 1/4th as long as apical anterior spur and subapical posterior spur; half as long as subapical anterior spurs. Hind legs with posterior spurs half as long as anterior spurs. Forewings: length 9.5 mm; membrane light-grey, with dark gray setae, without visible pattern, termen concave; first and second axillary sclerites enlarged on forewings and hind wings; R1 confluent with R2 shortly before C; base of discoidal cell located proximally of mid-length of wing; fork I longer than fork II; forks I and II present; fork I sessile, fork II short petiolate, almost sessile, crossveins s, r-m and m-cu present; postanal vein present. Hind wings: R1 meeting R2 apically; fork I slightly longer than fork II.

Male genitalia. Abdominal segment IX fused annularly, each side with 1 longitudinal groove running apicad, widening into sclerotized plate on apical lobes dorsally of coxopodites; dorsal longitudinal grooves absent, but with vestigial path forming fold in lateral and dorsal views; tergum slightly longer than venter; anterior margins vertical, nearly straight. Apicodorsal lobes rounded, extending beyond bases of preanal appendages. Lateroapical corners of tergum IX forming short, triangular plates over articulation of preanal appendages. Antecostae well developed, absent on dorsum, forming darkly pigmented marginal rims connected to ventral longitudinal grooves; spine rows absent on posterior margins of segment IX; apicopleural and apicoventral areas without setae. Segment IX nearly glabrous, with few setal alveoli scattered around apical lobes and on ventropleural region. Intersegmental depression between segment IX and segment X with small slope. Segment X weakly pigmented, quadrangular in lateral view; with rounded apex longer than gonocoxites; in dorsal view, central ridge separating lateral concavities delineated by 2 long, dark, parallel-sided, slightly narrowing, lines running to near apex of segment. Apicoventral setose lobes with few subapical setae, below glabrous concavities. Apicodorsal setose areas not discernible. Apex of segment X filled, dorsal interlobular gap minute, triangular. Preanal appendages long, filiform in lateral and dorsal view. Each coxopodite and harpago together slightly shorter than segment X and longer than preanal appendages; coxopodites straight, almost parallel-sided in lateral view; slightly curving in ventral view; harpagones narrowing apically, packed with small, cone-shaped setae in ventral view. Phallic apparatus 2-partite in lateral view. Sclerotized part of phallotheca with sigmoid dorsum in lateral view. Endotheca and phallicata indiscernible from each other. Phallotremal sclerite small, not clearly visible.

Holotype male: LAOS PDR: Luang Phrabang Prov. : Nam Sat stream, 50 m upstream Hoi Sat Village, 336 m, UTM 48Q0220672, 2260175, 28.iv–2.v.2005, Malaise trap, loc 16 [N. Jönsson, T. Malm & B. Viklund] – ( NRM).

Distribution: Laos.

Etymology: Jonssoni, named after Mr. Niklas Jönsson (NRM), one of the collectors of the species.

Figures

FIGURES 178–180. Marilia jonssoni, new species, holotype. 178 —head, frontal; 179 —head, dorsal; 180 — right wings.

FIGURES 181–184. Marilia jonssoni, new species, holotype. 181 — genitalia, lateral; 182 — genitalia, dorsal; 183 — gonocoxite, ventral; 184 — phallus, lateral.