Marilia sumatrana Ulmer, 1951

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 93-96

Marilia sumatrana Ulmer

Figs 211–216

Marilia sumatrana Ulmer, 1951: 11 .

Type locality: Indonesia (Sumatra).

New records: LAOS PDR: Luang Namtha Prov.: Nam Ha NBCA, Lakkhammai Village, Nam Leung stream, 749 m, UTM 47Q0744602, 2339873, 30.iv.2005, light trap, loc 29 [N. Jönsson, T. Malm & B. Viklund] – 3 males ( NRM) ; Tong Om Village , 552 m, UTM 47Q0750111, 2321825, 1.v.2005, light trap, loc 30 [N. Jönsson, T. Malm & B. Viklund] – 1 male ( NRM) . Vientiane Prov.: Vang Vieng, Nam Xong River , upstream bamboo footbridge, 363 m, UTM 48Q0223506, 2115465, 26.iv.2005, light trap, loc 10 [N. Jönsson, T. Malm & B. Viklund] – 1 male ( OPC) . VIETNAM: Bac Thai Province: Quang Chu , 24–25.v.1987, light [J. Oláh] – 5 males, 9 females ( OPC) . Cuc Phuong : 13.x.1963 [T. Pócs] – 1 male ( OPC) .

This whitish animal with snow-white abdomen and whitish-hyaline wings belongs to the diagnostic species-group characterized by having whitish colour on some of their body parts, i.e. M. lata Ulmer, 1926 , M. simulans Forsslund, 1936 ; M. javana Ulmer, 1951 , M. parallela Hwang, 1957 , and M. albofusca Schmid, 1959 . Marilia tuyetmira , new species is easily distinguished from the other species by the presence of a long, narrow and oblique lateroapical corner of tergum IX, and there is no complete anterior suture on tergum IX. The distance between the eyes in dorsal view is longer compared to in M. namha ; the vertexal medioantennal compact setal wart are separate, rounded; and the occipital setal warts are rounded triangular, not drop-shaped. Marilia tuyetmira is distinguished from M. lata as the latter species has a convex termen on the forewings; genitalia with broad, filiform preanal appendages; basally very broad coxopodites in ventral view; and the maxillary palp formula is I-(IV,V)-II-III. Marilia simulans is separated from M. tuyetmira in having a convex termen; the forewing venation is different, like the shorter M1+2; and segment X is more pointed in dorsal view. Marilia javana has curved coxopodites and very narrow segment X in dorsal view. Marilia parallela is distinguished by its forewing venation. Marilia albofusca is distinguished by genitalic characters, i.e. having ventral part of segment X more sclerotized, and with produced apicoventral corners

Marilia tuyetmira , new species

Figs 217–223

.

Male (in alcohol). Body medium-sized; brown; legs, antennae and palps slightly lighter than body; abdomen, cephalic and thoracic setal warts white; forewing membrane whitish hyaline. Eyes black; moderately large; vertex slightly constricted,eyes; distance between eyes equal to eye diameter, and 2 times longer than broad. Ocelli absent. Tentorium slender, with short, vestigial dorsal arm; posterior arms robust, ending in pair of large, posterior tentorial pits; tentorial bridge slender, without protuberances; posterior half of anterior tentorial arms very thin, anterior half more robust, with strongly developed median lamellate lobes. Facial groove pattern simple. Pair of short, narrow, brown, fragmented frontal lateral setose wart with white alveoli, and enlarged antennal sockets dominating on frons. Anterior part of frontal grooves absent. Frontogenal vertical grooves invisible. Frontogenal, fragmented warts long, brown, vertically elongated. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; running obliquely laterad, reaching subgenal process. Subantennal grooves nearly indiscernible; vertical, forming narrow strip between ocellar groove and frontogenal compact setose warts; subocular groove not visible. Vertexal groove pattern simple; coronal groove almost complete, forming straight line in middle of vertex; anterior part vestigial. Postoccipital grooves encircle foramen magnum and postoccipital lobes. Labrum forming small, short, triangular structure with median setal area; anterior part movable, freely hanging. Mandibles almost indiscernible; lacinia visible, broad, setose. Head dorsum with small, rounded vertexal lateroantennal compact setal warts. Vertexal mediantennal compact setal warts well separated, large, round. Occipital compact setose warts large; triangular at posterior end of coronal groove. Pair of sensilla basiconica rounded, located along end of coronial groove, near occipital warts. Pair of postgenal compact warts following shape of elongate postgenal area, curving along posterior part of ocular grooves; forming narrow strip near ocular groove. Postgenal surface glabrous. Maxillary palps light brown; maxillary palp formula V-IV-III-II-I; each basal segment without setose subapicomesal nodule. Each scapes shorter than head. Each pedicel shorter than first segment of each flagellum; flagellae almost 2 times longer than forewings. Pronotum with 1 pair white warts with white setae; transversely elongate, occupying entire segment, almost touching mesally, narrowest laterally. Mesoscutal diffuse pair of setose warts composed of few light alveoli arranged in 2 longitudinally elongate, short, narrow, irregular patch on middle of dorsal surface of mesoscutum, separated by white median notal groove. Almost entire surface of mesoscutellar area white, with few, nearly indiscernible alveoli; few diffuse warts scattered among numerous small dots. Each proepisternum with small-sized, ovoid setose wart, smaller than setose wart on precoxale. Large, compact, setal wart present anteriorly on cervical sclerites; forming sclerotized surface on membranous part of neck, touching only anterior cervical sclerite. Lateral cervical sclerites forming narrow anterior arm articulating anteriorly to back of head with occipital condyle above posterior tentorial pits, fused to posterior cervical sclerites. Posterior cervical sclerite forming posterad broadening plate reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerites; dark pattern of cervical sclerite complex clearly visible on pale membranous neck. Leg claws symmetrical; spur formula 2, 4, 4; foreleg spurs almost equally long; midleg posteroapical spurs 2 times longer than foreleg spurs; midleg posterior subapical spurs 1/4th as long as anterior spurs. Hind leg posterior spurs half as long as anterior spurs. Forewings: 8.0 mm; narrow, with concave termen; membrane whitish hyaline, with whitish setal pattern by groups of darker setae; R1 confluent with R2 shortly before C; base of discoidal cell located proximally of mid-length of wing; fork I longer than fork II; forks III–V absent; fork I sessile, fork II short petiolate; crossveins s, r-m and m-cu present; postanal vein present. Hind wings: R1 present, long, fusing with R2 near apicalmost part of C; fork I slightly longer than fork II; with long basal brush. First and second axillary sclerites enlarged on both forewings and hind wings.

Male genitalia. Abdominal segment IX fused annularly, with 1 longitudinal groove running apicad, not reaching distal margin of apical lobes; dorsal longitudinal grooves absent; tergum longer than venter; anterior margins convex, with 2 small projections; each dorsal projection triangular, located at end of antecostal sutures and ventrally at confluence of ventral sutures; posterior margins with flat apical lobe deeply excised at preanal appendages. Apicolateral corners of tergum IX forming short, narrow, oblique plate over articulation of preanal appendages. Antecostae well developed, lacking on dorsum, forming dark marginal rims connected to longitudinal grooves; spine rows absent on posterior margins of segment IX; setose areas absent on apicopleural and apicoventral areas. Intersegmental depression between segment IX and segment X slightly sloping. Segment X weakly pigmented, forming quadrangular plate with rounded, narrowing apex in lateral view; longer than gonocoxites; 2 central longitudinal ridges separating lateral concavities in dorsal view, running to subapical end of segment; pair of lateral ridges running to apicolateral depressions. Apicoventral setose lobes with few subapical setae. Apicodorsal setose area represented by few dorsal setae. Apex of segment X filled; dorsal interlobular gap minute. Preanal appendages long, filiform in lateral and dorsal view. Gonocoxites shorter than segment X; coxopodites straight, with curved dorsal margin, resulting in narrowing apical half in lateral view. Harpagones slightly clavate in ventral view, with small, cone-shaped setae. Phallic apparatus forming basally curving, subapically constricted phallotheca and membranous apical portion of endotheca. Phallicata indiscernible. Phallotremal sclerites small.

Holotype male: LAOS PDR: Vientiane Prov.: Vang Vieng, Nam Xong River , upstream bamboo footbridge, 363 m, UTM 48Q0223506, 2115465, 26.iv.2005, light trap, loc 10 [N. Jönsson, T. Malm & B. Viklund] – ( NRM).

Paratypes: same data as holotype - 1 male ( OPC) . Luang Namtha Prov.: Nam Ha NBCA, Lakkhammai Village, Nam Leung stream, 749 m, UTM47Q0744602, 2339873, 30.iv.2005, light trap, loc 29 [N. Jönsson, T. Malm & B. Viklund] – 1 male paratype ( NRM) . Odomxai Prov.: Odomxai Town, Nam Ko River , just upstr. bridge, 28.iv.2005, light trap, loc 21 [N. Jönsson, T. Malm & B. Viklund] – 1 male paratype ( NRM) .

Distribution: Laos.

Etymology: Tuyetmira, from “mira tuyet”, snowy in Vietnamese, referring to the snow-white colour of the abdomen.

Figures

FIGURES 211–212. Marilia sumatrana Ulmer, from Vietnam: Bac Thai Province, Quang Chu. 211 — male head, dorsal; 212 — female head, dorsal.

FIGURES 213–216. Marilia sumatrana Ulmer, from Vietnam: Bac Thai Province, Quang Chu. 213 — genitalia, lateral; 214 — genitalia, dorsal; 215 — gonocoxite, ventral; 216 — phallus, lateral.

FIGURES 217–219. Marilia tuyetmira, new species, holotype. 217 —head, frontal; 218 —head, dorsal; 219 — right forewing.

FIGURES 220–223. Marilia tuyetmira, new species, holotype. 220 — genitalia, lateral; 221 — genitalia, dorsal; 222 — gonocoxite, ventral; 223 — phallus, lateral.