Marilia katakaha, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457, Zootaxa 2457 (1), pp. 1-128 : 81-83

Marilia katakaha , new species

Figs 185–188

The extremely narrow vertex is similar to that of M. mendolonga , new species. Marilia katakaha is separated from M. mendolonga in being larger. In addition, the vertexal lateroantennal compact setal warts of M. katakaha are larger, more rounded, with lateral part more elevated and supplied with larger alveoli compared to on the lower mesal part; while those of M. mendolonga are smaller and extremely elongate. Marilia katakaha has a ridge pattern on segment X being more strongly developed, particularly the lateral longitudinal ridges running to the marginal ridges of the depression are more pronounced.

Male (in alcohol). Body medium-sized; light brown; legs, antennae and palps light brown; wings uniformly light brown, without pattern in alcohol. Eyes faded brown, extremely large. Vertex forming narrow strip; interocular distance about 1/10th the eye width, almost 12 times longer than broad. Ocelli absent. Tentorium slender, with short, filiform vestigial dorsal arm; posterior arms robust, ending in pair of large posterior tentorial pits; tentorial bridge separating anterior and posterior tentorial arms very slender, without anteromesal or posteromesal protuberance; posterior half of anterior tentorial arms thin; anterior half robust, with strongly developed median lamellate lobes. Facial groove pattern reduced; short, narrow, frontal lateral compact setose wart and narrow, long, frontogenal compact setose wart, together with large antennal sockets dominating on frons. Anterior remnant of frontal grooves branching from coronal grooves invisible at ventral margin of antennal sockets. Frontogenal vertical groove invisible. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; long, running obliquely laterad, reaching subgenal process. Subantennal grooves running vertically in narrow strips between ocellar grooves and frontogenal compact setose wart; subocular groove indiscernible, merging to anterior clypeogenal grooves. Vertex forming narrow strip. Vertexal groove pattern simple; coronal groove ending in deep posterior cleft. Postoccipital groove encircling foramen magnum forming pair of postoccipital setal lobes. Labrum elongate, with short setae on hanging, freely movable anterior apical area. Mandibles almost indiscernible; laciniae broad. Pair of short, narrow compact frontal lateral setal warts present on face beside elongate, narrow pair of frontogenal compact setal warts running from antennal sockets along subantennal groove of pregenae. Pair of large, rounded vertexal lateroantennal compact setal warts visible on broad anterior section of narrow strip of vertex; lateral part elevated, supplied with large alveoli compared to on lower mesal part. Pair of vertexal mediantennal compact setose warts separated by septa. Occipital compact setose warts small, triangular. Pair of sensilla basiconicae present along end of coronial grooves, isolated from anteromesal corner of occipital warts. Vertically elongated postgenal compact warts curving along posterior section of ocular grooves, narrow strip pressed to ocular groove. Postgenal surface glabrous, postgenal surface hidden by eyes. Maxillary palps broken, 1 paratype has single basal segment present, with no setose subapicomesal nodule. Each scape slightly shorter than head. Each pedicel shorter than first segment of each antennal flagella. One pair pronotal warts present, transversely elongate, located on posterior half of pronotum, almost touching mesally, narrowest laterally; lateral ends almost separated into lateral, second pair setal warts. One pair mesoscutal diffuse setose warts present, represented by few alveoli visible on brown mesoscutal surface. Nearly entire surface of mesoscutellar area covered by diffuse setal warts represented by few alveoli. Each proepisternum with ovoid setose wart much smaller than setose wart on precoxale. Large, compact, setal wart present anteriorly on cervical sclerites; apparently forming sclerotized surface on membranous part of neck, touching anterior cervical sclerite. Lateral cervical sclerites forming narrow anterior arm articulating anteriorly to back of head with occipital condyle above posterior tentorial pits, fused to posterior cervical sclerites. Posterior cervical sclerite forming posterad broadening plate reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerites; dark pattern of cervical sclerite complex clearly visible on pale membranous neck. Legs claws symmetrical; spur formula 2, 4, 4; foreleg spurs almost equal; midleg posterior subapical spurs half as long as foreleg spurs; midleg posteroapical spurs 1/4th as long as foreleg spurs; hind leg posterior spurs half as long as anterior spurs. Forewings: length 9.5 mm; forewing membrane light brown, without visible pattern, termen straight; R1 confluent with R2 shortly before C; base of discoidal cell located proximally of mid-length of wing; fork I longer than fork II; forks III–V absent; fork I sessile, fork II short petiolate, nearly sessile. Hind wings: R1 long, running to R2 near append of C; fork I slightly longer than fork II; long setae in brush present on basal lobe.

Male genitalia. Abdominal segment IX fused annularly, ventral longitudinal grooves pronounced, separating ventral and dorsal half in lateral view, running nearly to apex of apical lobes; tergum IX slightly longer than venter; anterior margins straight vertical, with small triangular projection at end of antecostal sutures; posterior margins with rounded apical lobes; with deep excision at preanal appendages. Lateroapical corners of tergum IX forming short, narrow, transverse plate over articulation of preanal appendages. Antecosta on anterior margins of segment IX well developed, forming dark marginal rims connected to longitudinal ventral grooves. Ventral longitudinal grooves turning ventrad, forming short posterior grooves on segment IX; spine rows absent on posterior margins of segment IX; setose areas absent on apicopleural and apicoventral areas. Intersegmental depression between segment IX and segment X sloping gently. Segment X weakly pigmented, in lateral view forming quadrangular plate with slightly excised apex, ventral corner longer than preanal appendages and gonocoxites; central ridge long, triangular in dorsal view; additional pair of lateral ridges running to lateral depression; lateral depression half as long as segment X. Apicoventral setose lobes with few alveoli on lateral lobes. Apicodorsal setose area not discernible. Apex of segment X excavated, with Vshaped, dorsal interlobular gap. Preanal appendages long, filiform, almost parallel-sided in lateral and dorsal view. Gonocoxites shorter than segment X; coxopodites curved, with apical half narrow in lateral and ventral view; harpagones small. Phallic apparatus with basal half of phallotheca sclerotized; apical half of endotheca membranous; phallicata present. Sclerotized phallotheca curving basally, slightly constricted before apex. Phallotremal sclerite small, elongate, located at start of phallicata, U-shaped in ventral view.

Holotype male: MALAYSIA: O. Borneo: Songri Boh. [Mjöberg] – ( NRM).

Paratypes: same data as holotype – 1 male ( NRM); O. Borneo, Long Navang [Mjöberg] – 2 male ( NRM) .

Distribution: Malaysia (Borneo).

Etymology: Katakaha, ridge in Sanskrit; named after the complex system of dorsal ridges on segment X.

Figures

FIGURES 185–188. Marilia katakaha, new species, holotype. 185 — genitalia, lateral; 186 — genitalia, dorsal; 187 — gonocoxite, ventral; 188 — phallus, lateral.