Magnolia allasoniae MARTINETTO, 2022

Niccolini, Gabriele, Martinetto, Edoardo, Lanini, Benedetta, Menichetti, Elena, Fusco, Fabio, Hakobyan, Elen & Bertini, Adele, 2022, Late Messinian Flora From The Post-Evaporitic Deposits Of The Piedmont Basin (Northwest Italy), Fossil Imprint 78 (1), pp. 189-216 : 198-199

Genus Magnolia L., 1753

Magnolia allasoniae MARTINETTO sp. nov. validation of the species formerly described by Martinetto (1995: Chronological framing etc., pp. 90–91, in Italian)

H o l o t y p e. MGPT-PU141081, figured by Martinetto (1995: pl. 1, fig. 5), black and white photograph (Textfig. 4a), and newly photographed for this paper (Textfig. 4a2–a4).

P a r a t y p e s. MGPT-PU141082 ( Text-fig. 4b 1–b 4 ), figured by Martinetto (1995: pl. 1, fig. 4); MGPT-PU141083 ( Text-fig. 4c 1–c 4 ), figured by Martinetto (1995: pl. 1, fig. 6); MGPT-PU141084 ( Text-fig. 4d 1–d 4 ), figured by Martinetto (1995: pl. 1, fig. 7).

P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.

PFN002957 (for new species).

R e g i s t r y n u m b e r. IFPNI ( Doweld 2016) registration record for the new species: C615F793-17A4- 68DD-56D5-7936C0A204D3.

R e p o s i t o r y. Presently, Museo di Geologia e Paleontologia dell’Università di Torino (acronym MGPT; see Pavia et al. 2017), Turin, Italy.

E t y m o l o g y. In honour of Dr. Barbara Allason, the first person who investigated the fossil (pollen) flora of Ca’ Viettone.

T y p e l o c a l i t y. Ca’ Viettone fossil site (45° 19′ N, 7° 37′ E), i.e., a series of outcrops along the Ca’ Viettone brook, close to the village of Levone Cavavese, Piemonte Region, Italy .

T y p e s t r a t u m. Silty clays of section 21 of the Ca’ Viettone locality (fig. 3.12 in Martinetto 1995), 15 m above the bed of the brook, on the right bank; named “Villafranchiano lithostratigraphic unit” by Martinetto et al. (2018), Zanclean. This informal name is still in use in the Piemonte region, solely for historical reasons, and designates the nonmarine deposits which conformably overlie the Pliocene marine successions .

D i a g n o s i s. Fossils consisting of the sclerotesta of seeds ( Manchester 1994), narrowly to broadly ovate or broadly obovate (heart shaped), mostly asymmetrical, dorsiventrally compressed, narrowly lenticular to narrowly ellipsoidal in cross-section, usually with a median longitudinal trough on the ventral side, mostly obtusely pointed at the micropilar end, rounded or truncated basally, without any bulge at the heteropyle, length 6.0 to 8.7 mm, length to width ratio of seeds 2.7–1.0: 1, length to thickness ratio 7.0–3.0: 1, thickness of sclerotesta 0.3– 0.5 mm, rugulate surface carrying through the inside of the sclerotesta, heteropyle located centrally, c- or v-shaped in ventral view, stalk oriented almost parallel to the long axis of seed or bent up to 20°.

D e s c r i p t i o n o f h o l o t y p e. Seed with a heartshaped outline and obtuse apex; size 8.2 × 7.0 mm; length to width ratio of seed 1.2: 1; external wall of the seed with a few, longitudinal and oblique, differently long round-edged ridges and furrows; chalaza pit 1.4 mm wide, heteropyle located centrally, v-shaped in ventral view, stalk oriented parallel to the long axis of seed.

R e m a r k s t o s p e c i e s v a l i d a t i o n. The work Martinetto (1995) represented the publication by Servizio EDSU (institution for the right of university studies of the Turin University) of 100 printed copies of a doctorate thesis, which were in part distributed to palaeobotanists around the world and in part remained available upon request. This publication can also be requested, since at least 30 years, at the Central National Library of Florence (https://opac.bncf.firenze.sbn.it/bncf-prod/resource?uri= TSI9700142&v=l&dcnr=7). Therefore, we suggest that such conditions fulfilled the ICN requirements for effective publication (see Art. 43.2., 43.3; Turland et al. 2018).

Martinetto (1995) introduced the new name M. allasoniae but failed to fulfill all the basic requirements of the International Code for Nomenclature (ICN; Turland et al. 2018) for effective publication: the collection hosting the holotype was not indicated (Art. 40.7 of the ICN). The name M. allasoniae was also cited in Bertoldi and Martinetto (1995), with an obscure reference to the collection hosting the holotype, but “a full and direct reference to the place of valid publication, with page or plate reference and date” (Art. 41.5 of the ICN; Turland et al. 2018) was not provided either there or in other publications, which indicates that the name M. allasoniae was not yet validly published. In order to validate the species ( Martinetto 1995), here we provide the information required by the ICN, including a diagnosis in English and a new description of the holotype (Art. 43.1, required after 1 January 1996), with terminology adapted from Manchester (1994). New photographs of the holotype and paratypes are also provided ( Text-fig. 4 ).

Figures

Text-fig. 4. Type specimens of Magnolia allasoniae MARTINETTO sp. nov. from the Pliocene locality Ca’ Viettone. a1–a4: Holotype (MGPT-PU141081) in different views, i.e., as originally figured in Martinetto (1995: pl. 1, fig. 5) in a black and white print (a1), in a new digital photograph in basal view (a2), ventral view (a3) and dorsal view (a4); b1–b4: Paratype MGPT-PU141082 in different views, i.e., as originally figured in Martinetto (1995: pl. 1, fig. 4) in a black and white print (b1), in a new digital photograph in basal view (b2), ventral view (b3) and dorsal view (b4); c1–c4: Paratype MGPT-PU141083 in different views, i.e., as originally figured in Martinetto (1995: pl. 1, fig. 6) in a black and white print (c1), in a new digital photograph in basal view (c2), ventral view (c3) and dorsal view (c4); d1–d4: Paratype MGPT-PU141084 in different views, i.e., as originally figured in Martinetto (1995: pl. 1, fig. 7) in a black and white print (d1), in a new digital photograph in basal view (d2), ventral view (d3) and internal view (d4). Scale bars 1 mm.

Text-fig. 1. Geographical location of the studied post-evaporitic sections (Piedmont Basin): Govone (1), Sioneri (2), Ciabòt Cagna (3) and Pollenzo (4).