Proctodrilus opisthoductus Zicsi, 1985

Höser, Norbert, 2018, Interrelation of Proctodrilus species (Oligochaeta: Lumbricidae) with lessivage and layering in European soil profiles, Zootaxa 4496 (1), pp. 96-110 : 100-101

Proctodrilus opisthoductus Zicsi, 1985

In total, 493 adult individuals were found at 24 localities.

• Joia Mare (RO), transect Crişul Alb floodplain. 46°18'N, 22°13'E. Floodplain channel. 1 ind. at 4 cm depth, under 2 cm cover of fresh high-water loam.

• Suplac (RO), transect Tîrnava Mică floodplain. 46°23'N, 24°30'E. Floodplain edge, foot of slope. 2 ind. in Aw on h3 Lts.

• Cetatea de Baltă (RO), transect Tîrnava Mică floodplain. 46°14'N, 24°08'E. Floodplain centre. Profile with bandlike layers, youngest bands up to 1 cm thick. 2 ind. at 3 cm depth, in h3 Ls3, under band of fresh highwater loam.

• Blaj (RO), Tîrnava Mică floodplain. 46°10'N, 23°54'E. Exterior foot of levee deposits. 2 ind. at 6 cm depth, in h2 Lts.

• Beteşti/Betfalva (RO), transect slope on Fâneaţa Îngustă. 46°18'N, 25°07'E. Centre of floor of steep-sided depression, head of valley. Inverse profile, several layers; end of alluvial fan. 2 ind. at 5–7 cm depth, in h5 Lts ( Höser 2016).

• Hoghilag (RO), transect Tîrnava Mare floodplain. 46°14'N, 24°38'E. Floodplain centre, floodplain edge. 35 ind. at 3–10 cm depth, in Aw on h2 Lts, where in places overlying a layer with P. tuberculatus ( Höser 2015) . Near river below fresh, sandy high-water loam.

• Alma (RO), four transects Tîrnava Mare floodplain. 46°12'N, 24°30'E. Two floodplain terraces near river. Profile in places with bandlike layers. 74 ind. at 3–30 cm depth, in Aw, h3 Ls3, Lts, h5 Lt; at floodplain edge over rust-mottled Go.

• Micăsasa (RO), Tîrnava Mare floodplain. 46°06'N, 24°08'E. Floodplain terrace closest to river. Profile with bandlike layers. 4 ind. at 6–10 cm depth, in layer h3 Ls3, in transition to rust-mottled Lts with clay skins.

• Valea Lungă (RO), transect Tîrnava Mare floodplain. 46°07'N, 24°01'E. Floodplain centre, floodplain edge, lower slope. Profile with several layers. 17 ind. at 24–30 cm depth, in h3 Lt, mostly in h3/4 Ls3, where concentrated at layer boundary to Sl. Clay skins present.

• Spini (RO), transect Mureş floodplain. 45°50'N, 23°06'E. Floodplain centre, floodplain edge. Close to river profile with bandlike layers. 22 ind. at 12–20 cm depth, in h3 Ls3 at layer boundary to Sl, at floodplain edge in h2 Lts.

• Săcămaş (RO), transect gorge. 45°54'N, 22°41'E. Profile with several layers. 17 ind. at 8–17 cm depth, in h2 Ls3. In same profile P. antipai in the layer above, P. tuberculatus in the layer below ( Fig. 1 ).

• Dobra (RO), transect Mureş floodplain. 45°55'N, 22°35'E. Floodplain terrace closest to river. 18 ind. at 6–10 cm depth, in heavily penetrated in places with roots h3 Ls3 on the underside of a covering of young highwater loam and up to 2 cm depth in Lts below this covering, therefore at the transition to Aw/aB.

• Pojoga (RO), two transects Mureş floodplain. 46°00'N, 22°21'E. Three floodplain terraces close to river. Profile with bandlike layers. 40 ind. at 5–13 cm depth, in h3 Ls3, in places above a layer containing P. tuberculatus .

• Vărădia de Mureş (RO), transect Mureş floodplain. 45°59'N, 22°08'E. Interior edge of floodplain terrace closest to river. Profile with several layers. 1 ind. at 10 cm depth, in Ls3.

• Nicolae Bălcescu (RO), transect Mureş floodplain. 46°01'N, 22°06'E. Floodplain terrace closest to river. Profile with bandlike layers. 15 ind. at 2–12 cm depth, in heavily matted with roots and slightly stony h3 Ls 3 of the Aw. Above a layer with P. tuberculatus .

• Bata (RO), four transects Mureş floodplain. 46°02'N, 22°03'E. Four floodplain terraces. Profile near river bandlike, followed by several layers. 177 ind. at 2–22 cm depth, in h2 Lts or Lt of the floodplain edge, mostly in h3 Ls3 of floodplain centre; concentrated at layer boundaries, also in sand-poor humic krotovinas in sandy layers, and at the edge of alluvial fans at foot of wooded slope. In places ( Höser 2000: Fig. 1 , 2015: Table 3) over a layer containing clay skins with P. tuberculatus .

• Odvoş (RO), transect Mureş floodplain. 46°05'N, 21°50'E. Floodplain channel, floodplain centre far from river. 4 ind. at 2–4 cm depth, in h3 Lt of the Aw. Clay skins present.

• Şoimoş (RO), transect Mureş floodplain. 46°06'N, 21°43'E. Profile near river with bandlike layers. 20 ind. at 2–9 cm depth, in slightly gravelly h5 Ls3/Ls4 of the Aw, partly above sand layer. Close to river above a layer with P. tuberculatus ( Fig. 1 and Höser 2015).

• Lipova (RO), transect Mureş floodplain. 46°06'N, 21°44'E. Close to river. 3 ind. at 4–5 cm depth, in h3 Ls3 of the Aw.

• Vladimirescu / Glogowatz (RO), transect Mureş floodplain. 46°08'N, 21°24'E. Three floodplain terraces. 10 ind. at 5–15 cm depth, in h3 Lts or Ls3, in floodplain channel in wet h3 Lt.

• Peştere (RO), transect Timiş floodplain. 45°32'N, 22°10'E. Several floodplain terraces, slope. 6 ind. at 4–7 cm depth in h2 Ls3 of the Aw close to river and 1 ind. at 4 cm depth in h3 Lts of the Aw of a plateau on the slope.

• Bărbăteşti (RO), transect Gilort floodplain. 44°52'N, 23°31'E. 1 ind. at 5 cm depth, in h3 Ls3.

• Şomărtin/Martinsberg (RO), transect slope on Pârâul Nou. 45°50'N, 24°38'E. Slightly angled upper slope. 6 ind. at 6–12 cm depth, in h3 Lts, at lower end of the Ah. P. tuberculatus in the layer below.

• Cârţa / Kerz (RO), two transects Olt floodplain. 45°47'N, 24°35'E. Two floodplain terraces closest to river. 14 ind. at 3–5 cm depth, in h2 Lts, Lu, Ls3, mostly in Aw.

Figures

FIGURE 1. Ecological species separation in the genus Proctodrilus: Soil profiles with two or three co-occurring species, each in a different layer. For further details see text. Bottom right: Generalized scheme of preferred occurrence of Proctodrilus species in the soil profile: close to layer boundaries and in the layer with finer particle size.