Pygommatius misamis Scarbrough & Hill

Scarbrough, A. G. & Hill, H., 2005, Pygommatius Scarbrough & Marascia, a valid ommatiine genus (Diptera: Asilidae) with new combinations and new species from the Philippines, Zootaxa 1055, pp. 1-22 : 14-16

Pygommatius misamis Scarbrough & Hill , sp. nov.

Figs. 21–29

Male. Brown with yellow legs, wide apical corners of segments yellow. Body length 8.6 mm; wing length 7.9 mm. Head: Mostly brown setose anteriorly. Face yellowish­gray tomentose with sparse, scattered, white setae; two rows of thin brown bristles; mystax with stout, yellowish­white bristles; FHWR 1.0:11.9. Palpus and proboscis brown. Antenna yellowish­brown to brown; FWLR 1.0:1.9. Frons rusty yellow tomentose. Four or five postocular brown bristles dorsally, longest bristle bent forward about 1/3 distance toward ocellar tubercle.

Thorax: Mostly brown. Mesonotum with postpronotum brownish­yellow, postalar callus yellow; tomentum mostly brown, narrow side gray, posterior yellow. Setae sparse, brown mostly; two dorsocentral and four lateral bristles present. Scutellum light yellowish­gray tomentose, sparse white setose with two brown marginal bristles. Mediotergite brown, anatergite much lighter, brownish­yellow. Pleuron brown, narrowly below halter yellow; three white tomentose stripes, alternating with gray; sparse setae and white bristles present.

Wing: Identical to apoticius . Cell m1WR 1.0:7.0:7.5.

Leg: Fore coxa and fore trochanter yellow; remaining coxae and trochanters brown; coxae sparsely to abundantly brown setose apically. Femora and tibiae yellow except for wide brown band on hind femur and apical 1/3 to 1/2 of hind tibia. Fore femur with three or four stout, light brownish­yellow ventral bristles on basal 1/3. Middle femur with setalike bristles, mostly yellow, one or two anteroventral setae bristly, brown, basal posteroventral seta stout. Hind femur with anteroventral bristle, brown, thin and short basally, much longer and thick apically; short, thin posteroventral setae present, two stout bristles posterobasally; HFWLR 1.0:5.4. Tibiae with unusually long, thin bristles, mostly brown; fore tibia with thin fringe of long, yellow setae laterally. Fore and middle tarsi with basal tarsomere yellow, narrow apex yellow; bristles thin, brown except one on fore tarsus.

Abdomen: Mostly brown to black, brown setose; wide apical corners and margins yellow; tergite 6 with apical margin broadly rounded. Tergite 8 tuberculate apically.

Sternites with short, oppressed, brown setae; sternites 3–5 with stout, erect, mostly yellow bristles; sternite 5 with two low tubercles medially; apical corner of sternite 7 with five or six long, flat, basally curved stout bristles.

Terminalia ( Figs. 21–25 ): Epandrium with dorsal branch slender, digitate; middle branch wide, apex wide, asymmetrically spatulate dorsally, oblique in lateral view, apex pointed; ventral branch narrow, strongly recurved. Gonostylus with narrow apex, bulbus base. Aedeagus wide apically, sheath caplike dorsally; ejaculatory apodeme unusually narrow, without lateral flange. Inner margin of gonocoxite oblique. Hypandrium with narrow, transverse, cluster of bristles.

Female. Differs from male as follows. Body length 8.1 mm; wing length 7.1 mm; FHWR 1.0:8.9; FWLR 1.0:2.0; m1WR 1.0:2.8:2.9; HFWLR 1.0:6.3. Legs: Fore and middle femora with ventral rows mostly or entirely brown setae, none especially bristly. Abdomen: Wide lateral margins of most tergites and sternites yellow; apical sternites with sparse, thin, erect, yellow and brown setae. Terminalia ( Figs. 26–29 ): Tergite 8 broadly membranous apically. Sternite 8 strongly triangular apically. Spermatheca long, curved basally. Genital fork narrow basally, wide sclerites laterally.

Specimens examined. PHILIPPINES: Holotype ɗ, Mindanao, Misamis Or. [ Misamis Oriental], Mt. Empagatao, 1050–1200 m, wild banana leaves, 19–30.iv.81, H. Torrevillas (BPBM #16,574). Paratypes, 1 Ψ, [Mindanao] Misamis Or. [ Misamis Oriental], Minalwang, 1050 m, 24.iii–4.iv. ’61, H. Torrevillas (BPBM); 1 ɗ, [Mindanao] Misamis Or. [ Misamis Oriental], Minalwang, 1050–1200 m, 5– 9.iv.’61 (BPBM); 1 Ψ, P.I. Misamis Or., Minubanan, 1050–1200 m, 5– 9.iv.’61, H. Torrevillas (BPBM); 2 Ψ, P. I. Misamis Or., Mt. Kibungol, 20 km SE of Gingoog, 700–800 m, 9–18.iv.1960, W. Torrevillas (BPBM); 1 Ψ, Philippines, Camarines Sur: Mt. Isarog; 500 m, 4.iv.1962, H.M. Torrevillas (BPBM); 1 Ψ, Philippines, Camarines Sur: Mt. Isarog; 500–600 m, 5.iv.1963, H.M. Torrevillas (BPBM); 1 Ψ, Philippines, Camarines Sur: Mt. Isarog; 21–22.iv.1963, H.M. Torrevillas (BPBM); 1 Ψ, P.I. Mindanao, Bukidnon, 1250 m, Mt. Katanglad 4–9.xii.1959, L.A. Quate (BPBM);1 Ψ, Limay, Bataan P.I. (BPBM).

Etymology. The species name, misamis [feminine], is a noun in apposition, referring to the type locality ( Misamis Oriental).

Remarks. Most similar to P. apoticius and P. hypnus (Oldroyd) , males of P. misamis are recognized by the stout bristles below the fore femur, the two low, moundlike tubercles on sternite 5 medially, produced median apical margin and the unusually stout, basally curved bristles along the apical corner of sternite 8, and the combined characters of the terminalia ( Figs. 21–29 ). It differs from P. iriga and P. montanus by the short flagellum and dark pronotum. The flagellum is much longer and yellowish, and the prothorax is yellow in the latter two species. Pygommatius misamis runs to P. hypnus in Oldroyd’s key (1972) but is distinguished by the characters above and the yellow fore coxa which is brown in the latter species. Females are further distinguished by the long spermatheca and unusually narrow base of the genital fork.

Figures

FIGURES 21 – 29. Pygommatius misamis Scarbrough, terminalia. Male, 21 – 23. Lateral, dorsal, and ventral views; 24. Gonostylus, lateral view; 25. Aedeagus lateral view. Female, 26 – 27. Dorsal and ventral views; 28. Spermatheca; 29. Genital fork.