Orthosyntexis thanti J. Gao, Engel, Shih, & T. Gao, 2021

Gao, Jia, Engel, Michael S., Shih, Chungkun, Ren, Dong & Gao, Taiping, 2021, A new genus of anaxyelid wood wasps from the mid-Cretaceous and the phylogeny of Anaxyelidae (Hymenoptera), Journal of Hymenoptera Research 86, pp. 151-169 : 151

Orthosyntexis thanti J. Gao, Engel, Shih, & T. Gao sp. nov.

Figs 4 , 5 , 6 and 7

Etymology.

The specific epithet honours 3rd Secretary General of the United Nations and Burmese diplomat U Thant (1909-1974) and his dedication to seeking peace within and between nations.

Diagnosis.

Antennal scape length to width ratio more than 3. Forewing 1Rs proclival to R. Mesofemur longer than mesotibia, metafemur nearly as long as metatibia; mesotibial apical spurs somewhat shortened and not narrowed apically.

Holotype.

Female, no. CNU-HYM-MA2015102 (Figs 4 , 5 and 6 ).

Paratype.

Female, no. CNU-HYM-MA2015103 (Fig. 7 ).

Description.

Holotype [paratype measurements given in parentheses]: Body about 8.61 [8.42] mm long in dorsal view, antenna 2.85 [2.62] mm long in ventral view; forewing about 5.64 [5.48] mm in length, maximum width 2.09 [1.67] mm; hind wing about 4.21 [4.42] mm in length.

Head moderately large, slightly narrower than thorax. Head 1.98 [1.77] mm wide and 1.22 [1.31] mm long, nearly quadrate. Compound eyes large and hemispherical; mandible weakly bent, with apical margin vertical (parallel to mandibular base) and with lowest tooth not that elongate (Figs 5B and 7D ); labial palpus with three palpomeres (Fig. 5D ); antenna with 16 flagellomeres, scape 0.55 [0.46] mm long, maximum width 0.17 [0.14] mm; pedicel 0.37 [0.41] mm long, maximum width 0.13 [0.13] mm; flagellomere I shorter than scape, 0.15 [0.11] mm wide, 0.25 [0.21] mm long; flagellomere II 0.15 [0.12] mm wide, 0.17 [0.15] mm long, as long as individual lengths of remaining flagellomeres (Figs 5C and 7C ).

Thorax wide, width across tegulae 1.59 [1.54] mm; pronotum short, with slightly developed anterior notch and prominent hind margin, and with mediolongitudinal furrow dorsally. Mesoscutum with longitudinal sulcus and notauli strongly impressed; mesoscutellum tapering to acute apex; ratio of lengths of prescutum, median longitudinal sulcus between notauli, and mesoscuto-mesoscutellar sulcus and mesoscutellum 2.3/1/5.3 [2.2/1/4.9], notauli close to mesoscutellum. Legs spindly, mesofemur longer than mesotibia. Metafemur length 1.69 [1.43] mm, nearly as long as metatibia (length 1.68 [1.50] mm), thick subapically, narrowed apically. Protibia and metatibia with only one apical spur (Figs 6B, D and 7G ), mesotibia with two apical spurs (Figs 6C and 7H ); all tarsomeres with stiff apical setae; basitarsi long but shorter than remaining tarsomeres combined; tarsomere V elongate but shorter than basitarsus; pretarsal claws long, with curved apices and each with a single preapical tooth (Fig. 6E ).

Abdomen only slightly narrower than mesothorax. Ovipositor strongly serrate apically, short, not protruding beyond abdominal tip, full length 2.67 [2.51] mm (Figs 6F and 7F ).

Forewing with dense microtrichia but no coloration pattern, except for slightly darkened costal area. C and R thick, costal area narrower than C and R widths. Pterostigma completely sclerotized; Sc absent; 1r-rs and 2rs-m absent; 2r-rs issuing from pterostigma at its basal 1/3; 1Rs short and slightly proclival to R, about 0.7 × as long as 1M, meeting 1M at right angle (Fig. 7E ). 2Rs+M distinct but short, 0.1 × as long as 1Rs+M (Fig. 7E ). 2r-rs slightly proclival. 2+3M 1.6 × [1.2 ×] as long as 4M. Cell 1mcu pentagonal, about 2.0 × [2.1 ×] as long as wide; cell 2mcu hexagonal, about 1.1 × [1.4 ×] as long as wide. 2Cu 2.1 × [1.6 ×] as long as 1Cu. 1cu-a reclival, nearly 0.8 × as long as 1Cu. 1m-cu nearly as long as 3Cu, 3Cu shorter than 4Cu. 2m-cu nearly 0.8 × [0.6 ×] as long as 4M and 1.2 × [1.1 ×] as long as 1m-cu.

Hind wing Sc absent. Cell r closed. 1Rs (about 0.36 [0.32] mm in length) longer than 1M (about 0.26 [0.30] mm in length). 1rs-m (about 0.23 [0.20] mm in length), with an angle at 2Rs and shorter than 1Rs. 1M straight; m-cu absent; 2M+Cu present (about 0.40 [0.33] mm in length), free abscissa of Cu and cu-a (about 0.40 [0.43] mm in length) developed, 1Cu and cu-a straight.

Figures

Figure 4. One photograph and two line drawings of Orthosyntexis thanti sp. nov., holotype (specimen CNU-HYM-MA 2015102) female A dorsal view as preserved B line drawing of dorsal view with forewings and hind wings artificially extended from body C line drawing of forewing and hind wing. Scale bars: 2 mm (A, B); 1 mm (C).

Figure 5. Photographs of Orthosyntexis thanti sp. nov. A habitus in ventral view B mandible in frontal view C part of antenna D labial palpus. Abbreviations: Fla 1 and Fla 2, flagellomeres I and II; Lp 1, Lp 2 and Lp 3, labial palpomeres I, II and III; Pe, pedicel; Sc, scape.

Figure 6. Photographs of Orthosyntexis thanti sp. nov. A hind wing hamuli (white arrows) B left protibial spur C right mesotibial spurs D left metatibial spur E right mesotarsal pretarsal claw F ovipositor sheath in lateroventral view. Scale bars: 0.1 mm (A-E); 0.5 mm (F).

Figure 7. Photographs of Orthosyntexis thanti sp. nov., paratype (specimen CNU-HYM-MA 2015103) female A habitus in dorsal view B habitus in ventral view C part of antenna D mandible in frontal view E left forewing F ovipositor in lateroventral view G protibial spur H mesotibial spurs. Scale bars: 1 mm (A, B); 0.2 mm (C); 0.1 mm (D-H). Abbreviations: Fla 1 and Fla 2, flagellomeres I and II; Pe, pedicel; Sc, scape.