Cancellocochlea coolongolook, Shea & Colgan & Stanisic, 2012

Shea, M., Colgan, D. J. & Stanisic, J., 2012, 3585, Zootaxa 3585, pp. 1-109 : 79

Cancellocochlea coolongolook n. sp.

( Figs 29G; 31G; 33G; 35G; 37G; 39C; 40C; 41B)

Etymology. For the hamlet of Coolongolook.

Diagnosis. Shell very small, light orange-brown, biconcave with depressed spire. Protoconch sculpture reticulate consisting of 19 prominent, widely spaced, narrow spiral cords and prominent curved, widely spaced radial ribs. Teleoconch sculpture of numerous, prominent, widely spaced, slightly sinuate, prosocline radial ribs. Ribs on body whorl 44–67 (mean 57). Umbilicus wide cup-shaped. Epiphallus shorter than penis, entering penis through a bud-like verge with a lateral slit. Penis with an expanded apical portion becoming tubular distally; no pilasters present.

Type material examined. Holotype. AM C.463435, Bachelor State Forest, south of Coolongolook , New South Wales, (32° 15´07.8” S, 152° 20´28.9” E), 9.v.2009, coll. J. Stanisic. Paratypes. AM C.466292, AM C.466293, same data as holotype .

Description. Shell very small, light orange-brown, biconcave with depressed spire. Whorls 3.50–3.75, tightly coiled, the last inflated and descending in front. Sutures strongly impressed. Shell diameter 2.77–3.25 mm (mean 2.91 mm), height 1.38–1.57 mm (mean 1.45 mm), H/D 0.49–0.52 (mean 0.51). Protoconch flat, of 1.25 whorls, diameter 0.40–0.53 mm. Protoconch sculpture reticulate consisting of 19 prominent, widely spaced, narrow spiral cords and prominent curved, widely spaced radial ribs. Teleoconch sculpture of numerous, prominent, widely spaced, slightly sinuate, prosocline radial ribs. Ribs on body whorl 44–67 (mean 57), width of interstices on the first teleoconch whorl equal to width of three to less than six ribs; on the penultimate whorl equal to width of four to less than six ribs; each rib with one, two or more periostracal blades. Interstitial sculpture of low prominent microradial ribs and low, weaker microspiral cords forming weak beads at their intersection; number of microradials between ribs on the first teleoconch whorl 6–9; on first quarter of body whorl 9–11. Aperture broadly ovately-lunate. Parietal callus prominent, transparent. Umbilicus wide cup-shaped, diameter 0.74–1.01 mm (mean 0.89 mm), D/U 2.98–3.74 (mean 3.27). Based on 8 measured adults.

Reproductive tract with ovotestis containing two clumps of alveoli, with more than two alveolar lobes per clump. Hermaphroditic duct crescent-shaped. Spermatheca with a large circular to oval bulb. Penial retractor muscle inserting at the junction of the penis and epiphallus. Epiphallus shorter than penis, entering penis through a verge. Penis with an expanded apical portion becoming tubular distally. Verge bud-like with a lateral slit. No pilasters present. Vagina shorter than penis. Atrium short.

Distribution and habitat. Known only from Bachelor State Forest, south of Coolongolook, NSW; found in wet sclerophyll/rainforest, living under logs.

Remarks. Cancellocochlea coolongolook n. sp. has protoconch sculpture (reticulate) that more closely resembles that of some species currently assigned to Planorbacochlea (viz. P. yessabahensis , P. nambucca ) than to Ca. heatherae n. sp and seems to indicate the presence of a presently unrecognised local clade. While the penial verge structure is similar to that of Comboynea boorganna it could equally as well associate the species with some of the Planorbacochlea species. A yet-to-be described species ( Ca. cf. coolongolook .: Figs 30A; 32A; 34A) with similar protoconch characteristics and gross shell morphology also occurs in Boorganna Nature Reserve on the Comboyne Plateau where it is sympatric with Co. boorganna .