Verbascum creticum (L.) Cav., in Elench. Pl. Horti Matr. 39. 1803.

Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani & Civeyrel, Laure, 2023, Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa, PhytoKeys 225, pp. 115-152 : 115

Verbascum creticum (L.) Cav., in Elench. Pl. Horti Matr. 39. 1803.

Celsia cretica ≡ Celsia cretica L., in Syst. Veg. ed. 13: 470. 1774. Type: [Italy]. Ponae Ital. [Lectotype, designated here: Herb. Linn. No. 774.3 (LINN)! (image available at: http://linnean-online.org/7174/); isolectotype: Linn. No. 774.4 (LINN)! (image available at http://linnean-online.org/7175/)]. (1)

Verbascum lyratum = Verbascum lyratum Lam., Encyclop. 4: 223. 1797, nom. illegit., non V. lyratum Pourret, Mém. Acad. Toulouse, ser. 1, 3: 332. 1788 (= V. chaixii Vill).

Ditoxia lyrata = Ditoxia lyrata Raf., in Précis Découv. Somiol.: 40. 1814, nom. illegit.

Celsia lyrata = Celsia lyrata (Lam.) G.Don, in Gen. Syst. 4: 499. 1837, nom. illegit.

Thapsandra cretica = Thapsandra cretica (L.) Griseb., in Spic. Fl. Rumel. 2: 40. 1844.

Celsia cavanillesii = Celsia cavanillesii Kunze in Flora 29: 698. 1846.

Verbascum creticum = Verbascum creticum (L.) Kuntze, in Revis. Gen. Pl.: 468. 1891, comb. nov. superfl.

Celsia balearica = Celsia balearica Gand. in Fl. Eur. 16: 112. 1889, nom. inval.

Celsia cretica balearica = Celsia cretica subsp. balearica Gand. ex Gand., in Nov. Consp. Fl. Eur.:345. 1910.

Lasiake lyratum = Lasiake lyratum (Lam.) Raf., Fl. Tellur. 4: 89. 1838. nom. illegit.

Celsia verbascifolia = Celsia verbascifolia R.Hern. ex J.J.Rodr., Fl. Menorca: 94. 1901, nom. nud.

Celsia cretica pallenscens = Celsia cretica f. pallenscens Maire, in Bull. Soc. Hist. Nat. Afrique N. 22: 308. 1931. Type: [Algeria]. Bône, pentes de l’Edough, grès, 600 m., 30 April 1930, R. Maire s.n (lectotype, designated here: MPU [MPU002670]!; isolectotype: MPU [MPU002671]!). [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.mpu002670] (2)

Notes.

(1) Linnaeus (1774: 470) described Celsia cretica in Species Plantarum as new nomen specificum legitimum. The diagnosis consists of " Cretica, C. fol. Inferioribus lyratis, superiofibus ovati samplexi caulibus " followed by a synonym, " Blattaria perennis cretica incana, foliis binis conjugatis " that was cited from Morison (1680: 488). The protologue also includes a morphological description " Folia pinnato-lyrata alterna: Superiora simpilicis; cordato-amplexicolia. Calyces serrati. Filamenta 4: duobus superioribus pilofis; infeririobus, laevibus antheris que majoribis. Corolla flavae fundo superiore maculis 2. ferrugineis ". However, the collection locality is not indicated. Seven years later, in his Supplementum Plantarum Systematis Vegetabilium, Linnaeus [son] (1781: 281) gave a detailed morphological description of Celsia cretica , along with the statement " Habitat in India, Creta " followed by the symbol “♂” indicating the gender of the particular organs or individuals within a composite plate, according to Simpson (2010). Moreover, another polynomial " Verbascum foliis radicalibus ovatis petiolatis, caulinis oblongis sessilimus serratis, subtus tomentosis " from Miller (1758: 182, ic. t. 273) were cited in synonymy.

Among the original material of the genus Celsia , preserved in Linnaean herbaria, three sheets relevant to Celsia cretica were found: the first (Herb. Linn. No. 774.3, image available at: http://linnean-online.org/7174/) is annotated at the bottom " celsia cretica " and “774.3” at above the right hand of the sheet by Linnaeus. The second specimen (Herb. Linn. No. 774.4, image available at: http://linnean-online.org/7175/) is annotated " cretica " at the base and “774.4” at above the right corner of the sheet. Both specimens are clearly showing the characters mentioned in the Linnaeus [son] (1781) diagnosis. After careful examination of the available collections and consideration of all elements in the protologue, the sheet No. 774.3 is the most complete and well conserved, and it is designated here as the lectotype of the name Verbascum creticum . However, the sheet No. 774.4 is selected here as the isolectotype.

(2) In the protologue of Celsia cretica f. pallenscens , Maire (1931) provided the following locality “Bòne, pentes du Mont Edough, 300-700 m" but they did not specify the name of the herbarium where the type material has been stored. We traced three sheets (MPU002670, MPU002671), preserved at Her. MPU which is in complete accord with the protologue. The specimen MPU002670 is here chosen as the lectotype for the name Celsia cretica f. pallenscens .