Hedinomorpha martensorum, Golovatch, 2021

Golovatch, Sergei I., 2021, Review of the millipede genus Hedinomorpha Verhoeff, 1934, with descriptions of three new species collected in China by Professor Jochen Martens (Diplopoda Polydesmida, Paradoxosomatidae), Zootaxa 4984 (1), pp. 368-384 : 371-373

publication ID	https://doi.org/ 10.11646/zootaxa.4984.1.27
publication LSID	lsid:zoobank.org:pub:7D854C7C-E117-4C20-90FA-5E83896609B4
DOI	https://doi.org/10.5281/zenodo.4927890
persistent identifier	https://treatment.plazi.org/id/2F528782-8C68-FFA1-FF71-82BDFE8EFC39
treatment provided by	Plazi
scientific name	Hedinomorpha martensorum
status	sp. nov.

Treatment

Hedinomorpha martensorum spec. nov.

Figs 1–15 View FIGURES 1–4 View FIGURES 5–10 View FIGURES 11–15

Type material. Holotype male and 1 male paratype ( SMF), China, Qinghai Prov., Lake Koko Nor, southern shore, near Heimahe , 3400 m a.s.l., 22.VI.1996, J. Martens leg.

Name. This is the second species of the genus devoted to Jochen Martens, this time to honour the entire Martens family: Jochen, the collector, his wife Beate, their daughters Charlotte and Friederike, and their son Felix. I have been privileged to enjoy their warm home, company and hospitality many times during my stays at and visits to Mainz.

Diagnosis. Differs from other species of the genus by the following combination of characters: colouration brown, devoid of a vivid colour pattern; paraterga poorly-developed, their lateral calluses smooth and delimited by a complete and distinct sulcus only dorsally; pleurosternal carinae present on segments 2–8, traceable as bulges until segment 18; and the peculiar shapes of the solenophore and its outgrowths (see also Key below).

Description (male). Length of both holo- and paratype ca 17–18 mm, width of midbody pro- and metazonae 1.6 and 1.8 mm, respectively. Colouration in alcohol mainly brown ( Figs 1–4 View FIGURES 1–4 ), devoid of a vivid colour pattern; most of head, paraterga, venter, legs and tip of epiproct lighter, yellow- or light brown; antennomeres 6 and 7 infuscate, dark brown, but tip of antennae pallid. Paratype generally a little lighter brown than holotype.

Clypeolabral region moderately setose, vertex with a few setae; epicranial suture thin, but evident ( Fig. 2 View FIGURES 1–4 ). Antennae moderately long and only slightly clavate ( Figs 1, 2 View FIGURES 1–4 ), extending behind metatergum 2 when stretched dorsally (male). In length, antennomere 2>3=4=5=6>>1=7. Interantennal isthmus about 1.5x as broad as diameter of antennal socket ( Figs 1, 2 View FIGURES 1–4 ). Tegument generally smooth and shining, only in places slightly rugulose above and below paraterga; surface below paraterga microgranulate; pro- and metazonae very delicately shagreened, strictures between them very faintly striolate; caudal margins of metaterga above a simple limbus densely and finely incised ( Figs 3, 4 View FIGURES 1–4 ). Tergal setae largely abraded or broken, few remaining ones thin and moderately long, ca 1/4–1/3 as long as metaterga; setation pattern traceable as 2+ 2 in a transverse fore (= pre-sulcus) row. In width, collum <head <segment 3=4 <2 = 5–16, body gently and gradually tapering thereafter. Paraterga poorly developed, largely set at 1/3 of metatergal height, pore-bearing ones considerably thicker than poreless ones in lateral view, broadly rounded and delimited by a narrow sulcus on collum, clearly drawn both anteriad and caudad, and somewhat projecting beyond both tergal margins only on segment 2, caudal corner clearly rounded, never sharp and never extended behind rear tergal margin thereafter ( Fig. 1 View FIGURES 1–4 ). Lateral calluses subhorizontal, smooth and narrow, but evident, delimited by a distinct and complete sulcus only dorsally, with a fainter sulcus in caudal 1/3 also ventrally only on pore-bearing segments. Ozopores lateral, almost invisible from above, each lying inside an ovoid pit at ca 1/4–1/5 off caudal margin. Stricture between pro- and metazona thin and shallow, nearly smooth. Transverse metatergal sulci evident, thin, simple, slightly sinuate medially, far from reaching the bases of paraterga, present on segments 5–18, in places slightly punctured. Axial line missing. Pleurosternal carinae granulate ridges with rounded caudal flaps increasingly developed on segments 2–7, abruptly reduced to small flaps on segment 8, carinae being increasingly reduced and retained only as small bulges towards segment 18 ( Fig. 1 View FIGURES 1–4 ). Epiproct ( Figs 4 View FIGURES 1–4 , 5 View FIGURES 5–10 ) as usual, rather long, faintly concave to subtruncate at tip, lateral pre-apical papillae small. Hypoproct ( Fig. 6 View FIGURES 5–10 ) high, semi-circular, caudal margin with 1+1 long setae borne on flat rounded knobs. Sterna moderately setose, cross-impressions shallow, transverse sulci being slightly deeper than axial ones, without modifications except for a small, roundly subtrapeziform, poorly concave and setose lobe between coxae 4 ( Figs 2 View FIGURES 1–4 , 8 View FIGURES 5–10 ). No tubercles near gonopod aperture. Legs moderately long and slender, likely incrassate in male compared to a still unknown female, midbody ones ca 1.2–1.3x as long as body height, densely setose, neither adenostyles nor laterally swollen prefemora, ventral brushes on prefemora and tarsi until two last leg-pairs. In length, femur> tarsus> coxa = prefemur = tibia> postfemur.

Gonopods ( Figs 1, 2 View FIGURES 1–4 , 9–15 View FIGURES 5–10 View FIGURES 11–15 ) largely suberect, in situ their distal parts coiled and in touch with each other; coxite subcylindrical, setose distoventrally, ca 2/3 as long as femorite (fe); prefemoral (= densely setose) part nearly half as long as fe; seminal groove (sg) usually running along fe at bottom of a shallow mesal groove/gutter (g) before moving onto a distinct flagelliform solenomere (sl) squeezed between a particularly strongly developed lamina lateralis (q) and a much smaller lamina medialis (j), and supported by a more complex postfemoral part; the latter forming a subtriangular, apical, postfemoral, lateral lobe (l) demarcated at base by a transverse lateral sulcus, and being largely represented by a long, slender, ribbon-shaped, loosely coiled, distally barbed and apically subacuminate solenophore branch (sph) supplied with a distinct spine (k) near midway and a rounded process/lobe (lo) in distal 2/3. No other outgrowths at base of l in addition to both main branches (sl and sph) except for a small tooth (t) distal to sl base.