Eritrichium echinocaryum (I.M.Johnst.) Y.S.Lian & J.Q.Wang

Hao, Jia-Chen, Ahmad, Latif, Zhou, Yu & Liu, Quan-Ru, 2017, The identity of Eritrichium heterocarpum Y. S. Lian & J. Q. Wang (Boraginaceae), Phytotaxa 303 (3), pp. 165-172 : 170

Eritrichium echinocaryum (I.M.Johnst.) Y.S.Lian & J.Q.Wang ( Wang et al. 1980: 46)

Basionym: — Hackelia echinocarya I.M.Johnst. (1940: 54) .

Nomen nudum: — Ertrichium deapiens I.M.Johnst.

Type: — CHINA. Yunnan: Dêqên County, C. W.Wang 70292 (Holotype: GH-00097407!; Isotype: PE-01338120!, WUK-0120956!, NAS-00213175!, IBSC-0536529!)

= Eritrichium heterocarpum Y.S.Lian & J.Q.Wang ( Wang et al. 1980: 45), syn. nov.

Type: — CHINA. Qinghai: Tongren County, 1964, Zh.D.Wei, Mai 499 (Holotype: WUK-0296907!).

Distribution and habitat: — China: Qinghai (Xining, Gonghe, Tongde), Yunnan (Shangri-la, Dêqên), Sichuan (Baiyu), Tibet (Markam); in open dry slope with rubble.

Phenology: —Flowering and fruiting from July to September.

Taxonomic notes: — Eritrichium echinocaryum is a member of E. set. Eritrichiastrium ser. Deflexa Y.S.Lian & J.Q.Wang ( Wang et al. 1980) according to Wang’s system ( Wang et al. 1980), classified on the annual or biennial herbs, much branched stems, linear-oblong blades and short pedicels. Eritrichium echinocaryum is more closely related to E. thymifolium because of its annual herbs, leaf blade oblanceolate to linear-oblong and nutlet heteromorphy. However, E. echinocaryum differs by its corollas shape (campanulate-rotate vs. campanulate-tubular) and diameter (4–5 mm vs. ca. 2 mm), faucal appendages bearing obvious glands (vs. no glands), surface feature of nutlets without wings (obvious verrucae and bristles vs. sparsely pubescent) and marginal glochids with bristles (vs. hairless). The main differences between E. echinocaryum and E. thymifolium are summarised in Table 1.

Additional specimens examined: — CHINA. Qinghai: Tongde County, Ho Ting-nung, Bruce Bartholomew & Mike Gilbert, 64 (QTPMB, PE), Y.H.Wu et al. 5810, 6453 (QTPMB); Xining County, Z.H.Zhang et al. 5608 (QTPMB); Gonghe County, collector ignotus 72138 (PE). Yunnan: Shangri-la County, Zhongdian Exped. 708 (PE). Sichuan: Baiyu County, Kham Exped. 10-0660 (PE). Tibet: Markam County, J.C.Hao 15578, 15579 (BNU). Location unknown, K.T.Fu 1272 (WUK).