Ogcodes Latreille, 1797

Winterton, Shaun L., 2012, Review of Australasian spider flies (Diptera, Acroceridae) with a revision of Panops Lamarck, ZooKeys 172, pp. 7-75 : 57-58

Ogcodes Latreille, 1797 Figs 3C63-64

Ogcodes Latreille, 1797: 154 - Schlinger 1960: 245; Schlinger and Jefferies 1989: 377. Type species, Musca gibbosa Linnaeus, by subsequent monotypy ( Latreille 1802: 432).

Oncodes Meigen 1822: 99 [emendation of Ogcodes Latreille] - White 1914: 69; Hardy 1921: 77, 1946: 66; Paramonov 1955: 23, 1957: 531.

Note.

Synonymy and usage restricted to Australasian region fauna only; see Schlinger (1960) for more exhaustive list.

Diagnosis.

Body length: 3.0-5.0 mm [male], 4.0-8.0 mm [female]. Body shape not arched, colouration black, yellow or white, non-metallic; head much smaller than thorax width, shape sub-spherical; postocular ridge and occiput rounded; two or three ocelli, anterior ocellus sometimes absent; posterior margin of eye rounded; eye apilose; position of antennae on head adjacent to mouthparts; eyes contiguous above antennal base, not contiguous below antennal base; palpus absent; proboscis apparently absent; flagellum shape stylate; apex with terminal setae (or single seta); antenotum not collar-like behind head; subscutellum enlarged; tibial spines absent; pulvilli present; wing hyaline, markings absent; costa ending near wing apex, costal margin straight; humeral crossvein absent; radial veins straight; R1 inflated or not inflated distally; pterostigma and cell r1 membranous, not ribbed; only two radial veins present, R2+3 absent, R4+5 not reaching wing margin; medial vein compliment with M1, M2 and M3 present, or two M veins present; discal cell weakly formed or absent; medial veins not reaching wing margin; cell m3 absent; CuA1 absent; CuA2 separate from A1, ending just before wing margin; crossvein 2r-m absent; wing microtrichia absent; anal lobe well developed; alula well developed; abdominal tergites smooth, rounded (rarely with tubercles in fossil species); abdomen shape greatly rounded, inflated.

Included species.

Ogcodes is a distinctive and cosmopolitan genus and the most species-rich in the family. Thirty-four species in two subgenera ( Ogcodes and Protogcodes Schlinger, 1960) are listed by Schlinger and Jefferies (1989) for the Australasian region.

Comments.

Ogcodes is in need of revision and no recent keys to species have been published for the region. The most recent revision of the genus was by Schlinger (1960), but there are many undescribed species in collections and a world revision of the genus is needed. Ogcodes is a derived genus with a typical globose body, relatively small head and reduced wing venation. Characters which differentiate Ogcodes from all other Acroceridae genera include antennae proximal to mouthparts, palpi absent, proboscis very short, almost all wing cells absent or poorly formed, eyes apilose and R2+3 absent.