Vargapupa oharai Páll-Gergely, 2015

Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András & Asami, Takahiro, 2015, Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae), Zootaxa 3937 (1), pp. 1-49 : 42-43

Vargapupa oharai Páll-Gergely , n. sp.

Fig. 8 C.

Diagnosis. A very large, slender turriform, brownish species with a well-developed and a weaker basal keel.

Description. Shell brownish corneous or yellowish, slender turriform, widest at its base; there are 10.25 (n=2) regularly increasing whorls; first whorl of the protoconch rather smooth, the second whorl is finely ribbed; suture deeper on the upper whorls, especially at the protoconch; the coarse riblets are regular on the whole shell; keel blunt, the ribs of the body whorl are visible here too, especially close to the end of the keel (close to the columellarparietal transition of the peristome); there are rather shallow sutures on both sides of the keel where it joins the shell surface of the last whorl; keel starts from the back side of the most basal point apertural rim, and ends on the ventrolateral side; keel gradually becomes lower towards its end; there is a second, lower keel between the main keel and the umbilicus; it also starts from the back side of the apertural rim; main keel visible inside the aperture as a shallow canal; aperture round with angled columellar-parietal and parietal-palatal transitions; the lower incision (basal part of the aperture) is the starting point of the keel; apertural rim thickened and very much reflexed.

Measurements (in mm). H: 14.7–16.1, D: 5.3–5.6 (n=2).

Differential diagnosis. See under Vargapupa biheli n. sp.

Material. Vietnam, Ninh Binh Province, Cuc Phuong National Park, Cave of Prehistoric Man, 237 m, 20°17.56874'N, 105°40.01229'E, leg. Ohara, K., Okubo, K., Otani, J. U., 16.11.2007., Holotype HNHM 98837, OK/2, JUO/2; same data, leg. Okubo, K. & Otani, J. U., 12.04.2007., JUO/1, PGB/1; Vietnam, Ninh Binh Province, Cuc Phuong National Park, excoll Jose Ahuir Galindo, PGB/1; Vietnam, Thanh Hóa Prov., Pu Luong N. P., surroundings of village Am, 20°28.185'N, 105°13.309'E, collected by native people, 0 5.10.2010., HE/1 paratype.

Type locality. Vietnam, Ninh Binh Province, Cuc Phuong National Park, Cave of Prehistoric Man, 146 m, 20°15.53843'N, 105°42.38950'E.

Etymology. Named after our friend, Kenji Ohara, who provided the material.

Distribution ( Fig. 15 ): The new species is known from two localities in Northern Vietnam. There is approximately 60 km between the two sites.

Remarks. In the Cuc Phuong National Park Vargapupa oharai n. sp. lives sympatrically with Pseudopomatias amoenus .

Figures

FIGURE 15. Distribution of Pseudopomatias and Vargapupa n. gen. species. Empty triangle, top up: Vargapupa biheli n. sp.; filled triangle, top down: Vargapupa oharai n. sp.; empty square: Pseudopomatias maasseni n. sp., open circle: P. sophiae n. sp., closed circle: P. linanprietoae n. sp.

FIGURE 8. A. Csomapupa luyorensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3377; B. Cs. grandis (Godwin- Austen, 1876), syntype, NHMUK 1903.7.1.3438; C. Vargapupa oharai n. sp., holotype, HNHM 98837; D. Vargapupa biheli n. sp., holotype, MNHN 2012 – 27020; E. Nodopomatias oakesi (Godwin-Austen, 1917), syntype, NHMUK 1903.7.1.3086; F. Nodopomatias sibbumensis (Godwin-Austen, 1917), syntype, NHMUK 1903.7. 1.3146. Photos: A – B, E, F: H. Taylor; C – D: B. Páll-Gergely. Scale bars = 5 mm; the smaller one refers to the photos of the whole shell, the larger one refers to the photos showing the lateral and basal side of the shells.