Pseudopomatias nitens Páll-Gergely, 2015

Páll-Gergely, Barna, Fehér, Zoltán, Hunyadi, András & Asami, Takahiro, 2015, Revision of the genus Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae), Zootaxa 3937 (1), pp. 1-49 : 39-40

Pseudopomatias nitens Páll-Gergely , n. sp.

Figs 4 , 5 A, 6A, 6E, 9A–E.

Diagnosis. A small to medium sized, glossy, rather turriform species with plicae inside the last whorl.

Description. Shell glossy, yellowish to brownish (often "rusty") in colour, rather turriform (sometimes spindle-shaped); the last or the penultimate whorl is the widest; 6.25–6.75 moderately to very much bulging whorls are separated by a rather deep suture which is even deeper between the upper whorls; protoconch consist of 1.5 whorls, the first half is finely tuberculated, glossy, the remaining one whorl is regularly and very finely ribbed; teleoconch regularly, finely ribbed, until the aperture; umbilicus slit-like, very narrow; most specimens with three plicae on the dorsolateral side of the body whorl (for description of the inner plicae see "Remarks on the shell characters" and Fig. 4 ); aperture rounded, with not of very slightly angled columellar-parietal transition and slightly angled parietal-palatal transition; apertural rim paler than the shell colour, it is thickened and very slightly reflexed; parietal callus slightly thickened; rim double, consists of a weaker inner and a stronger outer circle separated by a deep suture.

Measurements (in mm). H: 6.0–8.7, D: 2.8–3.3 (different Vietnamese samples, n=4); H: 8.3–8.9, D: 3.4–3.7 (shells from Changyang, n=4); H: 7.4 D: 3.4 (HNHM 98834, Sichuan, n=1).

Differential diagnosis. Pseudopomatias nitens n. sp. differs from P. sophiae n. sp. by the more corpulent, spindle-shaped shell, the lower and denser ribs, the thinner apertural margin and the presence of inner plicae above the umbilicus in most nitens specimens. P. linanprietoae n. sp. is lighter in colour, larger and has a thicker apertural margin. P. nitens n. sp. differs from P. amoenus by the smaller, glossy, spindle-shaped shell, more bulging whorls, stronger ribs, the absence of spiral lines between the ribs (sometimes present in P. amoenus ), the more slender, relatively smaller shell, the more regularly shaped aperture, thicker peristome and the presence of two to three palatal plicae above the umbilicus that are visible throughout the semi-transparent shell. See also Table 6 .

Material. Tonkin, coll. Denis 1945, MNHN 2012-27023/1 holotype, MNHN 2012-26931/31 paratypes and 2 juveniles (not paratypes); Muong Kong, Tonkin, NHMUK 1909.7.2.36/1 paratype; Long Ping, Tonkin, NHMUK 1909.7.2.40/1 paratype; Tonkin, Pac-Kha, probably leg. Messager, NHMUK 1909.7.2.24/1 paratype; Tonkin, Long Ping, probably leg. Messager, NHMUK 1909.7.2.25/1 paratype; Muong-Hum, Tonkin, E. R. Sykes coll., Acc. no. 1825, NHMUK 20130424/1 paratype; Tonkin, Muong-Hum, coll. Letellier 1949, MNHN 2012-26932/2 paratypes; Tonkin, coll. Denis 1945, MNHN 2012-26936/3 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012- 26935 and P. amoenus : MNHN 2012-27018); Tonkin, Coll. Denis 1945, MNHN 2012-26942/1 not paratype, very slender shell without plicae; Tonkin, Coll. Denis 1945, MNHN 2012-26944/4 paratypes and 2 juvenile shells (not paratypes, one of them is very large, possibly P. amoenus ); Tonkin, coll. Denis 1945, MNHN 2012-26946/14 paratypes (mixed sample with P. amoenus : MNHN 2012-26945); Tonkin, pris du Col de Nuages (2000 m altitude), Tonkin, Muong-Hum, between Muong-Hum and the Clouds Pass (Col de Nuages), near the latter one, more than 2000 m a.s.l., leg. Messager, MNHN 2012-27022/1 paratype (mixed sample with P. amoenus : MNHN 2012- 26992); leg. Messager, MNHN 2012-26953/7 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012- 26952); Tonkin, leg. Messager, MNHN 2012-26960/1 paratype, 2 damaged shells (not paratypes) (mixed sample with P. sophiae n. sp.: MNHN 2012-26959); Indochine, leg. Messager, MNHN 2012-26962/1 corroded shell without plicae (not paratype); Indochine, leg. Messager, MNHN 2012-26969/3 paratypes and one juvenile shell, which is not paratype (mixed sample with P. amoenus: Messager , MNHN 2012-26968 and P. sophiae n. sp. Messager, MNHN 2012-26970); Indochine, leg. Messager, MNHN 2012-26972/2 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012-26971); Indochine, leg. Messager, MNHN 2012-26974/1 paratype (mixed sample with P. sophiae n. sp.: MNHN 2012-26973); Indochine, leg. Messager, MNHN 2012-26978/3 shells without plicae, not paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012-26977); Indochine, leg. Messager, MNHN 2012-26979/3 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012-26980); Indochine, leg. Messager, MNHN 2012-26982/4 paratypes (mixed sample with P. sophiae n. sp.: MNHN 2012-26981); Tonkin, leg. Messager, MNHN 2012-26987/1 shell without plicae, not paratype (mixed sample with P. sophiae n. sp.: MNHN 2012-26988); Indochine, leg. Messager, MNHN 2012-27006/20 paratypes (mixed sample with P. amoenus : MNHN 2012-27005); Tonkin, Pakhé, leg. Messager, MNHN 2012-27011/7 paratypes (mixed sample with P. amoenus : MNHN 2012-27009 and P. sophiae n. sp.: MNHN 2012-27010); Tonkin, leg. Messager, MNHN 2012-27016/2 paratypes (mixed sample with P. amoenus : MNHN 2012-27015); China, Changyang, B. R. Lucas coll, Acc. no. 2351, NHMUK 20130425.1 (not paratype), NHMUK 20130425.2–4 (not paratypes); Changya, Hupei, China, coll. Boettger, excoll Schmacker 1893, SMF 45650/8 (not paratypes); China, Patung, Hupei, coll. Naegele, SMF 50008/2 (not paratypes; mixed sample with P. amoenus : SMF 342542); China, Changya, coll. S. H. Jaeckel, SMF 211185 /2 (not paratypes); Changya, Carl Bock coll, NHMUK 1894.4.24.94–7/4 (not paratypes); Changyang, J.J. MacAndrew coll, NHMUK 20130426/3 (not paratypes); Changyang, E. R. Sykes coll, Acc. no 1825, NHMUK 20130427/2 (not paratypes); Changyang, excoll Dr. W. Blume, Dr. Leo P. Rušnov coll, NHMW 109201 (not paratypes); Changyang, NHMW 83901/G/8273 (not paratype); China, Sichuan, Dujiangyan Shi, Taian Zhen, Qingchenghoushan, Cuiyinghu to upper station of Jinli cable station, 1273 m, 30°56.27110'N, 103°28.75198'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 0 9.07.2007., HNHM 98834 (not paratype), OK/1; China, Sichuan, Dujiangyan Shi, Taian Zhen, Qingchenghoushan, above lower station of Jinli cable station, 979 m, 30°55.62540'N, 103°29.08667'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 0 8.07.2007., OK/1 (not paratype, juvenile specimen, it can be this species or P. amoenus ); China, Sichuan, Emeishan Shi, Eshan Zhen, Emeishan, Wanniansi, 865 m, N29°34.83012', E103°22.97325', leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 14.07.2007., OK/1 (not paratype).

Type locality. Tonkin.

Etymology. We use Preston's manuscript name. The Latin nitens means shiny, glossy, glittering, bright, which is characteristic for P. n i t e ns n. sp.

Distribution ( Fig. 11 ): The new species is known from Northern Vietnam (only the border region with Yunnan), and the Chinese Sichuan and Hubei Provinces.

Remarks. Specimens from Changyang and Badong are taller, have more whorls, rather spindle-shaped shells, more reflexed peristome and more rounded aperture. Shells from Sichuan have tumid ("ovoid") shape and the two circles of the peristome are equally developed and well separated.

In museum collections Pseudopomatias nitens n. sp. was frequently in mixed lots with P. amoenus and P. sophiae n. sp.

Figures

FIGURE 4. Plicae of Pseudopomatias nitens n. sp. A part of the last whorl was missing, therefore the plicae were visible. A. all plicae (enumerated). B – D. the first two plicae forming a by-pass canal. The squamous surface is probably the result of old, corroded material.

FIGURE 5. SEM images of the protoconch (A – B) and operculum (C, E – F: outer side, D: inner side) of Pseudopomatias sp. A. P. nitens n. sp., HNHM 98834; B. P. amoenus Möllendorff 1885, China, Yunnan, Yiliang Xian, Jiuxiang Fengjingqu, 1818 m, 25 ° 04.27306 ' N, 103 ° 22.87032 ' E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 10.06. 2005., coll. PGB; C. P. amoenus, Indochine, leg. Messager, MNHN 2012 - 26983; D. P. amoenus, Tonkin, Muong-Hum, between Muong-Hum and the Clouds Pass (Col de Nuages), near the latter one, more than 2000 m a. s. l., leg. Messager, MNHN 2012 - 26997; E – F. P. e o s, Nantou Xian, Renai Xiang, Meifeng-Songgang road, around km 14, 2120 m asl., 24 ° 05.103 ' N 121 ° 10.257 ' E, leg. Hunyadi, 20.10. 2014.

FIGURE 11. Distribution of Pseudopomatias species. Filled circle: P. pe g ue ns i s (Theobald, 1864); open triangle, top up: P. nitens n. sp.; open circle: P. amoenus Möllendorff 1885 s. l.; empty square: P. e o s Pilsbry & Hirase 1905; script " Shan States ": Pseudopomatias shanensis n. sp. The meaning of the numbers: 1: Sandoway, 2: Pegu, 3: locality of P. amoenus near Jiuxiang Fengjingqu, 4: locality of P. amoenus near Kongshan Cun, 5: Northern Vietnamese and Chinese localities of P. amoenus and P. nitens n. sp., 6: Mau Son, That Khé and the new localities of " typical fulvus " (= P. amoenus), 7: Xuan Son National Park; 8: Cuc Phuong National Park, 9: Chengkou (type locality of Fargesia cornea), 10: locality near Mufu Cun (new locality of typical P. amoenus), 11: Badong (type locality of P. amoenus), 12: Changyang (locality of P. nitens n. sp.), 13: Hengshan (possible position of the type locality of P. amoenus var. tumidula), 14: Guiyan Xian (possible position of the type locality of P. amoenus var. tumidula), 15: Nanling Guojia (see Hu et al. 2004).