Wardianum triangulare ( Harrah, 1922 )

Dronen, Norman O. & Blend, Charles K., 2015, Updated keys to the genera in the subfamilies of Cyclocoelidae Stossich, 1902, including a reconsideration of species assignments, species keys and the proposal of a new genus in Szidatitreminae Dronen, 2007, Zootaxa 4053 (1), pp. 1-100 : 47-48

Wardianum triangulare ( Harrah, 1922) — Type species

Syn. Wardianum catoptrophori Dronen, 2007

Type host. Pectoral sandpiper, Caladris melanotos (Vieillot) (Syn. Tringa maculata Vieillot ) ( Charadriiformes : Scolopacidae ).

Type locality. Creston, Iowa, USA.

Additional hosts. Willet, Tringa semipalmata (Gmelin) (Syn. Catoptrophorus semipalmatus [Gmelin]) ( Charadriiformes : Scolopacidae ). Also reported from species of Actitis Illiger , Ereunetes Illiger and Schistrophorus (unknown bird genus)— Yamaguti (1971).

Additional locality. Galveston, Texas, Iowa and Oklahoma, USA—Yamaguti (1971).

Previously proposed synonym. Corpopyrum wilsoni ( Harrah, 1922) (= W. wilsoni [ Harrah, 1922])— Bashkirova (1950).

Remarks. Although originally described as Cyclocoelum triangularum Harrah, 1922 from specimen No. 21.88 in the Ward Collection at the University of Illinois, this species was transferred to Wardianum by Witenberg (1923, 1926). It was considered to be a synonym of Haematotrephus lanceolatum ( Wedl, 1858) by Macko & Freige (1960). Dronen (2007b) described W. catoptrophori from willets in the south-central USA and although it has the ovary removed anteriorly from the anterior margins of the testes, uterine loops that appear to be somewhat more extensive laterally, vitelline fields that extend more anteriorly, and a somewhat narrower egg than described for W. triangulare , the latter appears to have been described from a smaller, contracted specimen and we consider the two species to be synonymous. Dronen (2007b) pointed out that unlike most species in the genus, W. catoptrophori had a pharynx that was wider than long; however, in our opinion the usefulness of this characteristic in terms of distinguishing species in the genus is not well enough documented. Rudimentary oral sucker present—Harrah (1922) and Bashkirova (1950).

= Nοt present. 1Estimated frοm figure in οriginal descriptiοn.