Wardianum facioi Brenes & Arroyo, 1962

Dronen, Norman O. & Blend, Charles K., 2015, Updated keys to the genera in the subfamilies of Cyclocoelidae Stossich, 1902, including a reconsideration of species assignments, species keys and the proposal of a new genus in Szidatitreminae Dronen, 2007, Zootaxa 4053 (1), pp. 1-100 : 49

W. facioi ( Brenes & Arroyo, 1962) n. comb.

Type host. Northern jacana or jaçana, Jacana spinosa spinosa (Linnaeus) ( Charadriiformes : Jacanidae ).

Type locality. Proviencia de Puntarenas, Costa Rica.

Remarks. This species was originally described as Cyclocoelum (Haematotrephus) facioi Brenes & Arroyo, 1962 . It was transferred to Haematotrephus by Yamaguti (1971). It was originally described as having the vitelline fields as “joining in the posterior end” (= confluent posteriorly); however, the illustration ( Fig. 5 of Brenes & Arroyo 1962) appears to show that the vitelline fields are not confluent posteriorly. In this species the pretesticular ovary forms a triangle with the testes (Haematotrephinae), the genital pore is postpharyngeal, the vitelline fields appear to not be confluent posteriorly and the testes are side by side, placing this species in Wardianum . However, if the vitelline fields are confluent posteriorly as originally described, it would likely require the erection of a new genus in the subfamily to facilitate this species. No oral or ventral sucker described—Brenes & Arroyo (1962).

Figures

FIGURES 5 – 9. Diagrammatic representations of species representing the genera assigned to Haematotrephinae Dollfus, 1948. 5. Neohaematotrephus Kanev, Radev, & Fried, 2002 represented by Neohaematotrephus arayae Zamparo, Brooks, Causey & Rodriguez, 2003, adapted from Zamparo et al. (2003); 6. Harrahium Witenberg, 1926 represented by H. halli (Harrah, 1922), adapted from Harrah (1922); 7. Uvitellina Witenberg, 1923 represented by Uvitellina himantopi Dronen & Tkach, 2013, adapted from Dronen & Tkach (2013); 8. Wardianum Witenberg, 1923 represented by Wardianum catoptrophori Dronen, 2007, adapted from Dronen (2007 b); 9. Haematotrephus Stossich, 1902 represented by Haematotrephus limnodromi Dronen, Gardner & Jiménez, 2006, adapted from Dronen et al. (2006 b). Abbreviations: C, cecum; G, genital pore; O, ovary; P, pharynx; S, rudimentary oral sucker; T, testis; U, uterus; V, vitelline field.