Lispe orbitalis, Pont, 2019

Pont, Adrian C., 2019, Studies on the Australian Muscidae (Diptera). VIII. The genus Lispe Latreille, 1797, Zootaxa 4557 (1), pp. 1-232 : 130-133

Lispe orbitalis sp. nov.

( Figs 272–283 )

Diagnosis. In the group of species with a seta on posterior apical margin of hind coxa, the male of L. orbitalis can be immediately recognised by the very long and partly outcurved lower orbital seta ( Figs 273–274 ), by the very narrow gena ( Figs 272–273 ), and by the conformation of the hind basal tarsomere ( Figs 276–278 ).

Etymology. The name L. orbitalis refers to the very strong and partly outwardly directed lower orbital seta ( Fig. 274 ).

Type-Material Examined: Holotype ♂. AUSTRALIA: Northern Territory: Batten Point , 15.54S 136.32E, 30 km NE by E of Borroloola, 30.x.1975, M.S. Upton ( ANIC).

Description ♂ (♀ not known). Head ( Figs 273–274 ). Ground-colour black. Frons at broadest point 0.34, at lunula 0.23, of maximum head-width (frontal view). Fronto-orbital plate thinly silvery-grey pruinose below, thinly so above and subshining between orbital setae and vertex; parafacial, face, gena and most of occiput also silverygrey pruinose; frontal triangle inconspicuous, narrow, dark brown when viewed from below. Lower orbital seta strong, longer than upper orbital, directed outwards and backwards; with a line of frontal setulae mesad of the orbital setae. Parafacial very narrow, half as broad as postpedicel; setulose on almost entire length. Antenna black; postpedicel 2.8 times as long as broad, falling short of mouth-margin by 0.5 its length. Arista plumose, the hairing at widest point 2.6 times width of postpedicel. Vibrissa strong. Gena narrow, 0.1 of vertical eye-length; genal setae rather strong and dense. Palpus dark brown, little swollen apically ( Fig. 275 ).

Thorax. Ground-colour black. Scutum and scutellum rather thinly grey dusted, more densely so in posterior view; postpronotal lobe, notopleuron and pleura grey dusted. Viewed from behind, scutum thinly and irregularly grey dusted, with black areas around prst ia and prst sa, and broadly over post sa and postalar callus; medially with a dark brown line beginning at suture and running to tip of scutellum, broadening out behind and occupying over middle third of scutellum. Anterior spiracle yellowish. Acr setulae in 4 irregular rows at suture. Dc 2+3. 2 pprn. 1+1 ia. 1+1 sa, both strong. 1 strong and 1 weaker proepisternal setae. 1 strong proepimeral, slightly shorter than the strong proepisternal, with 4 adjacent setulae. Anepisternum without a short seta in upper anterior corner; posterior row with 3 strong and 3 weak setae. 3 strong kepst, lower one equidistant from the upper two. Meron bare below spiracle and above hind coxa. Scutellum with disc densely setulose, sides and ventral surface bare.

Legs. Black, except for knees and base of tibiae which are yellowish, fore tibia yellow on basal third; apical tarsomeres reddish on fore and mid legs. Fore coxa bare behind. Fore femur with a row of short spinules on av surface; pv row complete, the setae long and fine, in basal half twice as long as femoral depth; pd row complete. Fore tibia with the ventral setulae rather erect and spinulose; with a strong submedian p seta; with d, pd and pv apical setae. Fore tarsomeres rather short, without modifications. Mid femur on av surface with several short setae in basal half, continued as short spinules in apical half; pv surface with 6–7 longer fine setae in basal half, continued as spinules in apical half; basal half also with a row of short strong a setae, culminating in a strong seta at middle; 2 p preapicals. Mid tibia with 1 strong p seta; with strong ad, av, v, pv and p apicals, the ad apical a little before apex. Mid tarsomeres without modifications. Hind coxa with a seta on posterior apical margin. Hind femur on av and pv surfaces with short spinulose setulae along entire length; otherwise with 2–3 av and 3–4 longer pv, all in basal half; ad row complete; 1 d preapical. Hind tibia with 1 long ad, with the ground-setulae on this surface long and erect; 1 av seta; ad and d preapical setae long and strong; with 1 av apical seta ( Fig. 276 ). Hind tarsus with basal tarsomere short, shorter than 2nd tarsomere, swollen at base on p to v surfaces, and bearing a long dense brush of setulae on basal half of these surfaces; v surface also with fine curled setulae in basal half ( Figs 276–278 ).

Wing. Clear. Tegula black, basicosta orange to brown. Crossvein r-m placed slightly apicad of the point where vein R1 enters costa; dm-cu more or less straight, longer than apical section of vein CuA1. Vein M running straight towards wing-margin. Calypters white, margins creamy. Knob of haltere black.

Abdomen. Ground-colour black. Tergites grey dusted, tinged with yellow; with black markings as in Fig. 279 , these continuing down on to sides of tergites, especially broadly over hind-margin of tergite 4 and fore-margin of tergite 5. Epandrium grey dusted. Sternites 1–4 grey dusted, sternite 5 shining. Setae only present on tergite 5, with 1 pair of median and 1 of lateral discals, and 3–4 pairs of marginals, all very strong and erect. Sternites 2–4 with sparse setulae ( Fig. 280 ).

Ƌ terminalia. Epandrium separated from tergite 5 by syntergosternite 8, which has 2 spiracles; with a pair of exceptionally long setae, 1.5 as long as vertical length of epandrium; hardly produced lobe-like ventrally (Fig. 281). Tergite 6 absent. Sternite 6 present as 2 small elongate plates, each one attached to a ventral prolongation from the sternite, all withdrawn beneath sternite 5. Surstylus absent, i.e. fused to epandrium without trace (Fig. 281). Cercal plate divided dorsally and ventrally (Fig. 282). Hypandrium attached at two points: to a rod originating from wall of epandrium close to upper outer edge of cercal plate; and via a tiny apparent praegonite to lower anterior corner of epandrium. Phallic complex ( Fig. 283 ): hypandrium with several finger-like processes on ventral edge, meeting at posterior end with base of epiphallus; praegonite and postgonite present, the latter short; phallapodeme large, at mid-length joined to hypandrium by a small bridge; phallus long, tube-like, simple, juxta large, without spinules, with a pair of bifurcate and finger-like processes flanking it at extreme base.

Measurements. Wing-length, 4.0 mm. Body-length, 5.5 mm.

Biology. Not known.

Distribution. Known only from Australia (NT).

Figures

FIGURE 272. Lispe orbitalis sp. nov., ♂ habitus (holotype).

FIGURES 273–278. Lispe orbitalis sp. nov., ♂. 273. Head, lateral view. 274. Frons, dorsal view. 275. Palpus. 276. Tip of hind tibia and hind tarsomere 1, anterior view. 277. Hind tarsomeres 1–2, dorsal view. 278. Hind tarsomere 1, posterior view. adanterodorsal, av—anteroventral, d—dorsal, or—orbital. (Holotype.)

FIGURES 279–280. Lispe orbitalis sp. nov., ♂. 279. Abdomen, dorsal view. 280. Sternites 1–5. (Holotype.)

FIGURE 283. Lispe orbitalis sp. nov., ♂ phallic complex, lateral view. epi—epiphallus, pg—postgonite. (Holotype.)