Socotralia reitteri, Novák & Purchart, 2012

Novák, Vladimír & Purchart, Luboš, 2012, New species of the genus Socotralia and first record of the genus Alogista from Socotra Island (Coleoptera: Tenebrionidae: Alleculinae), Acta Entomologica Musei Nationalis Pragae 52, pp. 323-336 : 331-332

Socotralia reitteri sp. nov.

( Figs. 11–15 )

Type locality. Yemen, Socotra Island, Firmihin.

Type material. HOLOTYPE: J ( NMPC), labelled:‘ YEMEN, SOCOTRA Island / Firmihin, 400-500 m / N 12°28′27″, E 54°0′54″ / 6.-7.ii.2010 at light / L. Purchart & J. Vybíral lgt .’ PARATYPES: 3 JJ 5 ♀♀, same data as holotype (1 ♀ NMPC, 1 J 2 ♀♀ LPCB, 2 JJ 2 ♀♀ VNCP).

Description. Male (holotype). Habitus as in Fig. 11 . Body small, elongate, from pale reddish-brown to dark blackish-brown, with pale brown setation, BL 4.6 mm. Widest near elytral midlength; BL/EW 2.56.

Head ( Fig. 12 ). Small, reddish-brown, with long, pale brown setation and microgranulation, slightly shiny. Anterior aspect and clypeus distinctly paler, punctation conspicuous, punctures relatively large. HW 0.9 mm; HW/PW 0.64; HL 0.5 mm. Eyes dark, large, transverse, deeply excised, space between eyes distinctly broader than antennomere 2 is long; OI equal to 22.28.

Antennae ( Fig. 13 ). Long, with microgranulation, punctation and pale brown setation, bicoloured, with antennomeres 1–2 and base of antennomere 3 pale reddish-brown, slightly shiny, anterior half of antennomere 3 and antennomeres 4–11 black, duller.AL 3.2 mm, AL/BL 0.70. Antennomere 2 shortest, antennomere 3–10 distinctly broader at apex. RLA (1–11): 0.77: 0.51: 1.00: 1.22: 1.20: 1.24: 1.20: 1.33: 1.31: 1.31: 1.45. RL/WA (1–11): 1.93: 1.56: 2.58: 2.46: 2.38: 2.38: 2.61: 2.51: 2.69: 2.64: 3.50.

Maxillary palpus. Pale reddish-brown with pale brown setation and microgranulation. Palpomeres 2–4 distinctly broadened from base to apex, slightly shiny. Ultimate palpomere broadly triangular.

Pronotum ( Fig. 12 ) unicoloured reddish-brown, slightly longer than semicircular, bellshaped, with long, pale brown setation, setae directed backwards, with microgranulation and punctation. Punctures conspicuous, relatively large. PL 1.0 mm; PW 1.4 mm; PI equal to 70.43. Borders complete, pronotal base bisinuate, on antescutellar area straight. Posterior angles rectangular, anterior angles indistinct.

Ventral part. Reddish-brown, with pale brown setation, microgranulation and punctation, punctures large and coarse.

Elytra unicoloured dark blackish-brown, elongate, covered by dense, long, backward directed, pale brown setation. EL 3.2 mm. Broadest near elytral two thirds, EW 1.8 mm; EL/ EW 1.78. Elytra with microgranulation and punctation; punctures relatively large. Rows of punctures in elytral striae not clearly conspicuous. Scutellum triangular, pale reddish-brown with pale brown setae and sparse microgranulation, shiny. Elytral epipleura well developed, pale reddish-brown, with large punctures and pale brown setation, regularly narrowing from elytral base to abdominal ventrite 1, then continues as parallel.

Legs. Unicoloured pale reddish-brown, with dense, short, pale brown setation. Tibiae, tarsi narrow, tibiae slightly dilated anteriorly. Penultimate tarsomere of each tarsus very slightly broadened, distinctly lobed. RLT: protarsus: 1.00: 0.72: 0.72: 0.80: 1.66; mesotarsus: 1.00: 0.44: 0.36: 0.48: 0.91; metatarsus: 1.00: 0.33: 0.30: 0.54. Both anterior tarsal claws with eight visible teeth.

Aedeagus ( Figs 14, 15 ). Pale brown, slightly shiny. Basal part rounded in lateral view, in dorsal view regularly narrowing anteriorly. Apical part (apex) short, in lateral view narrowly triangular, in dorsal view in form of a long triangle. AEB/AEA 3.54.

Female. More oval, antennae distinctly shorter in than male. Anterior tarsal claws with five visible teeth.

RLA (1–11): 0.71: 0.40: 1.00: 1.11: 1.09: 1.22: 1.31: 1.36: 1.27: 1.22: 1.29. RL/WA (1–11): 1.95: 1.00: 2.29: 2.26: 1.94: 2.03: 2.06: 2.21: 1.76: 1.91: 2.09.

RLT: protarsus: 1.00: 0.62: 0.76: 0.97: 1.76; mesotarsus: 1.00: 0.49: 0.44: 0.52: 0.88; metatarsus: 1.00: 0.35: 0.32: 0.48.

BL 4.4 mm; HL 0.5 mm; HW 0.8 mm; OI 29.86; PL 0.8 mm; PW 1.2 mm; PI 66.67; EL 3.2 mm; EW 1.7 mm; HW/PW 0.67; BL/EW 2.59; EL/EW 1.88; AL 2.6 mm; AL/BL 0.59.

Variability. Measurements: mean (minimum–maximum). Males (n=4). BL 4.5 mm (4.1–4.9 mm); HL 0.5 mm (0.4–0.6 mm); HW 0.8 mm (0.7–0.9 mm); OI 25.44 (22.28–28.85), PL 0.9 mm (0.8–1.0 mm) PW 1.2 mm (1.1–1.4 mm); PI 70.39 (68.61–71.59); EL 3.1 mm (2.8–3.5 mm); EW 1.7 mm (1.4–1.9 mm). Females (n=6). BL 4.3 mm (4.00– 4.5 mm); HL 0.4 mm (0.4–0.5 mm); HW 0.75 mm (0.7–0.8 mm); OI 33.10 (29.86–36.25), PL 0.8 mm (0.7–0.8 mm) PW 1.2 mm (1.1–1.3 mm); PI 66.89 (64.31–69.36); EL 3.1 mm (2.9–3.2 mm); EW 1.7 mm (1.5–1.9 mm).

Differential diagnosis (for more details see the key below). Socotralia reitteri sp. nov. differs from similar species S. montana sp. nov., S. vybirali sp. nov. and S. brunnea mainly in having antennomere 11 distinctly longer than antennomere 3, while S. montana sp. nov., S. vybirali sp. nov. and S. brunnea have antennomere 11 shorter than antennomere 3. S. reitteri sp. nov. differs from the similar species S. intermedia sp. nov. mainly in having space between eyes broader than length of antennomere 2, while S. intermedia sp. nov. has space between eyes distinctly shorter than length of antennomere 2. S. reitteri sp. nov. differs from the similar species S. major in having antennomere 3–10 distinctly serrate, while S. major has antennomeres 3–10 narrower. S. reitteri sp. nov. differs from the similar species S. minor in having sides of pronotum conically narrowing, while S. minor has rounded sides of pronotum.

Etymology. Named in honour of Edmund Reitter (1845–1920), world famous expert in Coleoptera .

Collecting circumstances. Members of this species were collected only at night when they were attracted to light. They were not found during the day on herb or shrub vegetation.

Distribution. Yemen: Socotra Island.

Figures

Figs. 11–20. Male holotypes of Socotralia reitteri sp. nov. (11–15) and S. vybirali sp. nov. (16–20). 11, 16 – habitus in dorsal view; 12, 17 – head and pronotum in dorsal view; 13, 18 – antenna; 14, 19 – aedeagus in lateral view; 15, 20 – aedeagus in dorsal view. Scale bars = 2 mm (body); 1 mm (antenna); 0.5 mm (pronotum).