Gomphrena filaginoides Martens & Galeotti (1843: 350)

Sandoval-Ortega, Manuel Higinio & Zumaya-Mendoza, Silvia, 2023, Identities of some names in Gomphrena (Amaranthaceae), Phytotaxa 626 (2), pp. 78-92 : 79

1. Gomphrena filaginoides Martens & Galeotti (1843: 350) ≡ Xeraea filaginoides (Martens & Galeotti) Kuntze (1891:

545).

Lectotype (here designated):— MEXICO, Oaxaca, dans les champs de la Misteca Alta et de la cordillère de Yavezia, 6000-7500 ft., April 1840, Galeotti 441 ( BR0000006950835 !, image available at https://www.botanicalcollections.be/specimen/ BR0000006950835 ); isolectotype P00622630!, image available at https://science.mnhn.fr/institution/mnhn/collection/p/item/p00622630).

= Gomphrena decumbens Jacquin (1804: 41–42) var. nana Stuchlík (1912: 158) ≡ Gomphrena nana (Stuchlík) Standley (1917: 150) .

Lectotype (here designated):— MEXICO, Manzanillo , 1 to 31 December 1890, Palmer 911 (Z000000292!, image available at https:// www.digitalis.uzh.ch/media/specimen/0/Z-000000292.jpg; isolectotypes: US00102786!, image available at http://n 2t.net/ark:/ 65665/30cd25a65-9c6c-4a31-8bd1-102acb23fdd6; US 00931477! [two specimens on the left side of the sheet], image available at http://n 2t.net/ark:/65665/3a3cde325-f174-4245-93b6-b0e8377de159).

Typification of the name Gomphrena filaginoides : — Martens & Galeotti (1843: 350) provided, after the description, the provenance (“dans les champs de la Misteca Alta et de la cordillère de Yavezia, de 6,000 à 7,000 piedes”); the citation “ Coll. H. Gal. N ° 441 bis” (which refers to Galeotti’s collection) is also given. We traced two specimens part of Galeotti’s collection (original material), one deposited at P (barcode P00622630) and another at BR (barcode BR0000006950835 ); both of these specimens match the protologue. BR0000006950835 ( Fig. 1 ) is here designated as lectotype; P00622630 is an isolectotype. Note that Galeotti’s collection and types are mainly preserved at BR (HUHIndex of Botanists 2023) .

Typification of the name Gomphrena decumbens var. nana :— The protologue of Gomphrena decumbens var. nana ( Stuchlík 1912: 158) consists of a brief morphological description, the citation of a collector and collection number (“Palmer 911”), as well as the mention of three herbaria, i.e. “Berl., Petr., Zür” (respectively, B, LE, and Z). One specimen is located in Z (barcode Z000000292), whereas duplicates were found at US (barcodes US00102786 and US00931477). The specimen Z000000292 ( Fig. 2 ) is here designated as lectotype being well preserved, showing mature reproductive structures, and matching the protologue .

Moquin-Tandon (1849: 417) includes Gomphrena filaginoides in the section “ species non tatis notae ” (= species not sufficiently known). Standley (1917: 153) cites G. filaginoides as a doubtful species and stated that it could be G. nana . Later, Holzhammer (1956: 196) mentioned that, according to the diagnosis and the original material (Palmer 911), the habit of G. decumbens var. nana is not distinct from G. filaginoides ; likewise, he reported the inflorescences of “ nana ” as coinciding with those of the type material of G. filaginoides . After reviewing the type material of G. filaginoides ( Fig. 1 ) and G. decumbens var. nana ( Fig. 2 ), we agree with Holzhammer (1956) and consider G. decumbens var. nana as a heterotypic synonym of G. filaginoides .

Figures

FIGURE 1. Lectotype of Gomphrena filaginoides (BR0000006950835).

FIGURE 2. Lectotype of Gomphrena decumbens var. nana (Z000000292 by United Herbaria Z+ZT / CC BY 4.0).