Ziminella, Korshunova, Tatiana, Martynov, Alexander, Bakken, Torkild, Evertsen, Jussi, Fletcher, Karin, Mudianta, I Wayan, Saito, Hiroshi, Lundin, Kennet, Michael Schroedl, & Picton, Bernard, 2017

Korshunova, Tatiana, Martynov, Alexander, Bakken, Torkild, Evertsen, Jussi, Fletcher, Karin, Mudianta, I Wayan, Saito, Hiroshi, Lundin, Kennet, Michael Schroedl, & Picton, Bernard, 2017, Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda), ZooKeys 717, pp. 1-139 : 16

Ziminella gen. n. Figs 2, 12, 13, 14, 15

Type species.

Eolis salmonacea Couthouy, 1838

Etymology.

In honour of Olga Zimina, scientist at Murmansk Marine Biology Institute; she made a considerable contribution in collecting Arctic paracoryphellid species for this study.

Diagnosis.

Body wide. Notal edge present, well-defined, continuous. Cerata not stalked, continuous. Rhinophores smooth to wrinkled, similar in size to oral tentacles. Anterior foot corners present. Anus pleuroproctic under the notal edge. Rachidian teeth with strong denticulated cusp; lateral denticles not clearly delineated from cusp. Lateral teeth weakly denticulated to smooth without attenuated process basally, significantly smaller than rachidian teeth. Receptaculum seminis not evident. Long vas deferens without separate granulated prostate. No penial collar. Penis folded or elongated conical.

Species included.

Ziminella abyssa sp. n. (Fig. 12), Ziminella circapolaris sp. n. (Fig. 13), Ziminella japonica (Volodchenko, 1941), comb. n. (Fig. 14) (original description in Volodchenko 1941; lectotype designated in Martynov 2013), Ziminella salmonacea (Couthouy, 1838), comb. n. (Fig. 15) (original description in Couthouy 1838; redescription in Kuzirian 1979).

Remarks.

Ziminella considerably differs morphologically from the genus Chlamylla by the absence of a granulated prostate and external penial collar (Fig. 7) and by the shape of the radular teeth, from the genus Paracoryphella by the presence of a penial sheath and by the shape of the radular teeth, from the genus Polaria by the absence of a very long, conspicuous distal receptaculum seminis and by the shape of the radular teeth. On the molecular tree, both species of the genus Ziminella place as a well-separated clade within the family Paracoryphellidae (Figs 1, 2). Four species currently known within the genus Ziminella are well-separated by the penial morphology ( Z. salmonacea possesses a slightly folded penis, whereas Z. abyssa and Z. japonica have an entire conical penis; for discussion see Martynov 2013). Molecular data (Figs 1, 2) published here for the first time show that three species are placed together within a larger clade.