Heinzia variabilis Korge, 1971

Gusarov, Vladimir I. & Koval, Alexander G., 2002, A revision of the genus Heinzia Korge, 1971 (Coleoptera: Staphylinidae: Quediina), with description of a new species and its probable larva, Zootaxa 69, pp. 1-19 : 11-12

2. Heinzia variabilis Korge, 1971 ( Figs. 9, 11 , 15, 17, 19 , 22­23 , 26­27 , 32 , 34 )

Heinzia variabilis Korge, 1971: 33 .

Heinzia variabilis: Coiffait, 1978: 284 . Heinzia variabilis: Herman, 2001: 3054 .

Type material. Holotype:, Turkey: Kaçkar­Da{lar near Iliça, valley of Ardesen (Ayder) 1000­1600 m (Korge & Heinz) (2.viii.1965) ( HKCB). Paratype:, the same data as the holotype ( HKCB).

Additional material. Turkey:, Palandöken­Da{, S of Erzerum, 2200 m, under stone near a creek bank (Heinz), 27.vii.1967 ( HKCB);, Riza, valley of Firtina, 1400 m (Vít), vii.1976 ( ASCO);, Trabzon, Kalkanl Da{lar, NW slopes, 8 km E of Zigana Pass, 1900­2100 m, alpine zone (A.Solodovnikov), 9.vi.1999 ( ASCC).

Diagnosis. Heinzia variabilis is very similar to H. caucasica and can be most reliably distinguished from the latter by less dense pronotal punctation and extensive impunctate areas of the disc ( Figs. 11, 10 ). Additionally, in H. variabilis impunctate area on the head is more extensive, reaching the center of the disc ( Fig. 9 ) (in H. caucasica the impunctate area is restricted mostly to anterior margin of the disc ( Fig. 8 )). Heinzia variabilis also differs from H. caucasica in having sharper subapical tooth of median lobe of aedeagus ( Figs. 32, 28­31 ) and longer medial process of female tergum 10 ( Figs. 34, 33 ).

Discussion. Number and position of punctures in dorsal rows and distribution of scattered punctation on the disc of pronotum varies in both H. variabilis ( Korge 1971) and H. caucasica . However, there is a significant gap in overall density of punctation between the two species ( Figs. 10­11 ).

Distribution. Heinzia variabilis is known from North­Eastern Turkey (altitude of 1000­2200 m) ( Fig. 48 ).

Natural History. One of the specimens of H. variabilis was found under stone near a creek ( Korge 1971). Another specimen (from Zigana Pass) was collected under deeply immersed stone at a creek bank (Solodovnikov, personal communication).

Figures

FIGURE 48. Geographical distribution of Heinzia Korge. Solid circles – H. caucasica Gusarov & Koval, sp. nov.; open circles – H. variabilis Korge.

FIGURES 8 ­ 13. Details of Heinzia caucasica Gusarov & Koval, sp. nov. (8, 10, holotype; 12 ­ 13, paratype from Baribana cave) and H. variabilis Korge (9, Kalkanl Da {lar; 11, holotype). 8 ­ 9 – head; 10 ­ 11 – pronotum; 12 – left antenna; 13 – right metatarsus. Scale bar 1 mm (8 ­ 12), 0.4 mm (13).

FIGURES 14 ­ 19. Details of Heinzia caucasica Gusarov & Koval, sp. nov. (14, 16, 18, paratype from Dolgaya cave) and H. variabilis Korge (15, 17, 19, Kalkanl Da {lar). 14 ­ 15 – male sternum 8; 16 ­ 17 – apex of mail sternum 8; 18 ­ 19 – male sternum 9. Scale bar 1 mm (14 ­ 15), 0.5 mm (16 ­ 19).

FIGURES 20 ­ 23. Aedeagus of Heinzia caucasica Gusarov & Koval, sp. nov. (20 ­ 21, paratype from Mezmay) and H. variabilis Korge (22 ­ 23, Kalkanl Da {lar). 20, 22 – ventral view; 21, 23, lateral view. Scale bar 0.4 mm.

FIGURES 24 ­ 27. Details of aedeagus of Heinzia caucasica Gusarov & Koval, sp. nov. (24 ­ 25, holotype) and H. variabilis Korge (26 ­ 27, holotype). 24, 26 – median lobe, ventral view; 25, 27, paramere. Scale bar 0.2 mm.

FIGURES 28 ­ 32. Apex of median lobe of aedeagus (lateral view) of Heinzia caucasica Gusarov & Koval, sp. nov. (28, paratypes from Mezmay; 29 ­ 30, Baribana cave; 31, Akhty) and H. variabilis Korge (32, Kalkanl Da {lar). Scale bar 0.2 mm.

FIGURES 33 ­ 35. Details of female tergum 10 of Heinzia caucasica Gusarov & Koval, sp. nov. (33, 35, Baribana cave) and H. variabilis Korge (34, paratype). 33 ­ 34 – dorsal view; 35 – lateral view; LL – lateral lobe of female tergum 10; MP – medial process of female tergum 10. Scale bar 0.3 mm.