Cratocerus monilicornis Dejean, 1829

Grzymala, Traci L. & Will, Kipling W., 2014, Taxonomic review of Cratocerus Dejean, 1829 (Coleoptera, Carabidae) with the description of six new species, ZooKeys 416, pp. 77-112 : 81-82

Taxon classification Animalia Coleoptera Carabidae

Cratocerus monilicornis Dejean, 1829 Figs 1, 9a, 11c

Cratocerus monilicornis Dejean 1829: 14-15; Dejean and Boisduval 1834: 17; Hope 1838: 84; Lacordaire 1854: 263-264; Gemminger and Harold 1868: 250; Chaudoir 1872: 18; Csiki 1929: 493; Blackwelder 1944: 34; Straneo 1977: 115; Lorenz 2005: 248.

Material.

1 male, labeled: "Museum Paris Montagnes des Orgues Prov de Rio-Janeiro Env. De la Tijuca Massif du Bico Do Papagaio E.R. Wagner 1902 700 m alt. Dicembre" (EMEC654896) borrowed from MNHN; 1 male, labeled: "Brazil, [Santa Catarina] Nova Teutonia, 27°11'S, 52°23'W, [-27.183 -52.383], Fritz Plaumann, 4.xi.1939" (no EMEC# assigned for this specimen, examined at MNSM); 1 female, labeled: "Corcovado Brazil GB 27-IV-1962 J. & B. Bechyne leg." (EMEC654898) borrowed from CUIC; 1 female, labeled: "Argt. Misiones Bemberg 29-1-45 Hayward-Wilink 60 bach" (EMEC654897) borrowed from MNHN.

Diagnosis.

This species is easily distinguished from all other congeners by its relatively large overall body size (9.5-12.0 mm), two pairs of supraorbital setae, and rounded lateral lobes of the mentum.

Description.

Size large for the genus, ABL 9.5-12.0 mm, TW 4.0-5.0 mm. Color. Head, clypeus, mandibles, pronotum, elytra, and venter rufo-piceous; labrum, antennae, remaining mouthparts, and legs fusco-ferrugineous. Head. HL 1.6-1.9 mm, HW 1.4-1.7 mm. Two supraorbital setae present over each eye. Apices of mandibles slightly curved, apices at less than 45 dedgree angle to mandibular midline. Apical maxillary palpomere length 5 × width at base. Mentum with rounded lateral lobes. Thorax. Pronotum (Fig. 9a) with distinct median impression, one distinct basal fovea laterad; one lateral seta located on apicolateral pronotal bead; one posterior seta present on posterior bead; pronotal anterior margin convex, greatest width at center; pronotal width 1.3-1.4 × pronotal length. Elytral length 1.4-1.5 × combined elytral width. Legs. Protibia with five distinct, stout spines confined to distal half on lateral margin. Genitalia, male. Median lobe curved sharply, between forty-five and ninety degrees, with respect to basal 1/3, remaining 2/3 with continuing curvature; not expanded apically; length average for genus. Left paramere complexly curved (Figure 11c). Genitalia, female. Spermatheca not examined for this species.

Distribution.

This species is geographically distributed in Brazil and northeastern Argentina (Fig. 13). According to Csiki 1929 and Blackwelder 1944, Cratocerus monilicornis has also been recorded from Venezuela, but specimens from this locality were not examined during this study.

Additional literature.

Dejean and Boisduval 1832: fig. 3; Ragusa 1887: 210.