Teratomyza formosana, Papp, László, 2011

Papp, László, 2011, Oriental Teratomyzidae (Diptera: Schizophora), Zootaxa 2916, pp. 1-34 : 8-10

Teratomyza formosana sp. nov.

( Figs 12–13 , 21–27 , 30–31 , 111 ) Type material. Holotype 3, TAIWAN: Ilan Hsien, Fu-Shan LTER Site, Sep 27, 2000, along/over a small brook, leg. L. Papp, No. 11 (HNHM). Paratypes (all HNHM): 1 3: same as for holotype; 2 3 1 Ƥ (without head): ibid., along/over a small creek, No. 10, March 27, 2003, leg. L. Papp; 13: ibid., along/over a small river, No. 4, March 26; 3 3 2 Ƥ (1 male and 1 female gen. prep.): ibid., small creeks and partly dry rocky beds, March 26, No. 6.

Description. Measurements in mm: body length 1.62 (holotype), 1.70–2.00 (paratype 3), 2.41–2.50 (paratype Ƥ); wing length 2.05 (holotype), 2.20–2.40 (paratype 3), 2.64–2.81 (paratype Ƥ); wing width 0.70 (holotype), 0.75–0.78 (paratype 3), 0.85–0.87 (paratype Ƥ). Length of arista/antenna: 32/18.

Head. Eyes with dense ommatrichia. Frons with greasy shine. Inner vertical setae particularly long, 0.30–0.35 mm. Genal setae sparse, longest anterior seta 0.15 mm (holotype), only 2–3 setae behind (other than postgenal setae), which are thin and only 1 of them is longer than the others (up to 0.11 mm). First flagellomere large, discoid, slightly decumbent in some specimens. First flagellomere covered by rather long (0.015 mm) whitish cilia. Arista 0.35–0.37 mm.

Thorax. Mesonotum with a weak shine, at the same time its coverage of fine grey microtrichia is obvious. Postscutellum distinct.

Wing membrane unicolorous light brown, veins slightly darker ( Fig. 111 ). Length of costal sections (all wing measurements in mm) between R1 and R2+3 1.13, between R2+3 and R4+5 0.53, between R4+5 and M1+2 0.265, intercrossvein section 0.075, costal seta 0.08, length of basal cell 0.08, length of discal cell 0.275, distance between apices of M veins 0.58. Costal vein continued on a section of 0.02 mm over the apex of R4+5. Vein A1 almost straight. Female wing margin sections: R2+3–R4+5–M1–M4–A1: 52–53/27–28/63/38–39. Halter dark brown, stalk short (c. 0.10 mm), knob not clearly separable from stalk, 0.19–0.23 mm long but only 0.05–0.055 mm broad, tapering apically.

Legs almost unicolorous light brown. Pulvilli white, lateral appendages not particularly dense.

Abdomen. Preabdominal tergites without peculiarities, dark brown, anterior margins lighter, male sternite 5 quadrate. Sclerites medial to synsclerite 7 less sclerotised, not distinct ( Fig. 21 , cf. Fig. 9 ). Male genitalia. Subepandrial sclerite ( Fig. 12 ) with broader dorsal part. Cerci ( Fig. 12 ) with long setae, particularly towards apex. Surstylus ( Fig. 13 ) shorter than in T. chinica , apical part broader and only an apical quarter is without setae. Pregonite ( Fig. 24 ) without narrow apical process. Postgonite ( Fig. 26 ) much longer than that of T. chinica . Epiphallus ( Fig. 27 ) very short, shorter than broad at base. Mesophallus short as in T. chinica but the ribbons are half as broad. Phallapodeme ( Fig. 25 ) very short, only slightly longer than broad when viewed dorsally. Ejaculatory apodeme ( Figs 22–23 ) with narrow blade.

Female sternite 8 ( Fig. 30 ) rather pentagonal than hexagonal though cranial margin rounded. Spermathecae ( Fig. 31 ) more pear shaped (length/width 0.075 * 0.055 mm), all surface covered by large warts.

Differentiating characters are given in the key below.

Etymology. The specific epithet refers to the old (Portuguese) name of Taiwan, ‘Formosa’.

Figures

FIGURES 9 – 13. Teratomyza spp., male postabdomen and genitalia. 9 – 11 = Teratomyza chinica Yang Chikun, 199 [8]: 9 = pregenital sclerites, incl. syntergosternite, ventral view, 10 = epandrium, surstyli, cerci and subepandrial sclerite (dotted), subventral (anterior) view, 11 = surstylus, broadest extension (sublateral view). 12 – 13 = T. formosana sp. n.: 12 = epandrium, surstyli, cerci and subepandrial sclerite (dotted), subventral (anterior) view, 13 = surstylus, broadest extension (sublateral view). Scales: 0.2 mm for Figs 9 – 10, 12, 0.1 mm for Figs 11, 13.

FIGURES 21 – 27. Teratomyza formosana sp. n., male postabdomen and genitalia. 21 = pregenital sclerites, incl. syntergosternite, ventral view, 22 = ejaculatory apodeme, lateral view, 23 = same, dorsal view, 24 = pregonite, broadest extension (nearly lateral view), 25 = phallapodeme, dorsal view, 26 = postgonite, lateral view, 27 = epiphallus, caudal view. Scales: 0.2 mm for Figs 21 – 22, 0.1 mm for Figs 23 – 27.

FIGURES 28 – 31. Teratomyza species, female genitalia. 28 – 29 = T. chinica Yang Chikun: 28 = sternite 8, 29 = spermatheca. 30 – 31 = T. formosana sp. n.: 30 = sternite 8, 31 = spermatheca. Scale: 0.1 mm.

FIGURES 111 – 113. Wings of Teratomyzidae new species. 111 = Teratomyza formosana sp. nov.; 112 = Poecilovitila barbata sp. nov.; 113 = P. brevicornis sp. nov. Scales: 1.0 mm each.