Myopsocidae, Pearman, 1936
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publication ID |
https://doi.org/ 10.11646/zootaxa.5377.1.1 |
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publication LSID |
lsid:zoobank.org:pub:2BA30329-4BDF-4595-9864-3CA23D56A659 |
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DOI |
https://doi.org/10.5281/zenodo.10248779 |
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persistent identifier |
https://treatment.plazi.org/id/345F87BE-FB0A-FFF6-D9C8-38D6AFBAFEBC |
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treatment provided by |
Plazi |
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scientific name |
Myopsocidae |
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Key to South American species of Myopsocidae View in CoL
( M. cinereus Navas, 1932 , not included because there are no illustrations or photographs in the original description)
1. Hindwing with Rs and M veins fused for a distance ( Fig. 68 View FIGURES 67–72 ).................................................... Myopsocus View in CoL .......................................................................................... 2
- Hindwing with Rs and M veins joined by a transverse vein ( Fig. 2 View FIGURES 1–5 )............................................... Lichenomima View in CoL ....................................................................................... 25
2. Species in which only females are known.................................................................. 3
- Species in which males and females or only males are known.................................................. 7
3. Forewings festooned apically, mainly on the apical margin of cell m2 ( Fig. 79 View FIGURES 79–84 ).................................... 4
- Forewings not festooned apically ( Fig. 67 View FIGURES 67–72 )................................................................. 6
4. Valve V1 basally narrower, with a dorsal tooth-shaped projection............................ M. vespertilio Mockford View in CoL
- Valve V1 with uniform width, without dorsal projections...................................................... 5
5. Forewing with large areas of depigmentation on M1 and M2 cells............................ M. albomaculatus View in CoL (New)
- Forewing with or without small depigmented areas M1 and M2 cells; pterostigma darker in distal half..... M. cinctus View in CoL (New)
6. Apical sclerite process subgenital plate long, narrow and with setae......................... M. albus (Garcia-Aldrete) View in CoL
- Apical sclerite process subgenital plate very short and without setae........................... M. circularis Mockford View in CoL
7. Forewings festooned apically, especially on the apical margin of cell M2. ( Figs 73 View FIGURES 73–78 , 79 View FIGURES 79–84 , 85 View FIGURES 85–90 )........................... 8
- Forewings without festooned apical margin ( Fig. 133 View FIGURES 133–138 )....................................................... 12
8. Hypandrium with latero-posterior processes ( Figs 77 View FIGURES 73–78 , 113 View FIGURES 109–114 ).................................................... 9
- Hypandrium without latero-posterior-processes ( Figs 83 View FIGURES 79–84 , 119 View FIGURES 115–120 )................................................. 10
9. Hypandrium widened apically, with widely separated latero-posterior processes ( Fig. 113 View FIGURES 109–114 ); phallosome with side struts convergent basally ( Fig. 114 View FIGURES 109–114 )........................................................... M. colombianus n. sp.
- Hypandrium narrow apically, with latero-posterior processes close together ( Fig. 77 View FIGURES 73–78 ); phallosome with side strut well separated basally, external parameres and side struts more widened than in the previous one ( Fig. 78 View FIGURES 73–78 ).......... M. amazonensis n. sp.
10. Hypandrium V-shaped, apically widened ( Fig. 119 View FIGURES 115–120 ); phallosome with side struts basally fused, very narrow proximally ( Fig. 120 View FIGURES 115–120 ), external parameres broadly rounded apically.......................................... M. garcialdretei n. sp.
- Hypandrium V-shaped, apically narrow ( Fig. 83 View FIGURES 79–84 ); phallosome with side struts basally separated but medially fused, external parameres not widened apically as above................................................................. 11
11. Forewing with M1 and M2 cells with apical hyaline area reaching to the wing margin ( Fig. 79 View FIGURES 79–84 ); endophallic membrane without spicules ( Fig. 84 View FIGURES 79–84 )...................................................................... M. andaquies n. sp.
- Forewing without hyaline areas in cells M1 and M2 as above ( Fig. 127 View FIGURES 127–132 ); endophallic membrane with abundant spicules ( Fig. 132 View FIGURES 127–132 )...................................................................................... M. inga n. sp.
12. Epiproct with conspicuous posterior horn.................................................. M. dispar (Enderlein) View in CoL
- Epiproct without posterior horn ( Fig. 136 View FIGURES 133–138 )................................................................. 13
13. Hypandrium subtrapezoidal, posterior margin with small tubercles on both sides of a narrow medial process ( Fig. 137 View FIGURES 133–138 ).......................................................................................... M. insularis n. sp.
- Hypandrium with variable shape, without small tubercles and medial process as described above ( Figs 95 View FIGURES 91–96 , 107 View FIGURES 103–108 , 168 View FIGURES 163–169 )..... 14
14. Clunium male with a pair of rounded lobes on the dorsum, covered by small spicules and setae ( Fig. 178 View FIGURES 170–180 ); phallosome oval ( Fig. 177 View FIGURES 170–180 ).............................................................................. M. minutus (Mockford) View in CoL
- Clunium male without a pair of rounded lobes on the dorsum; phallosome with variable shape, sometimes oval ( Figs 96 View FIGURES 91–96 , 108 View FIGURES 103–108 , 150 View FIGURES 145–150 , 169 View FIGURES 163–169 )........................................................................................... 15
15. Hypandrium with latero-posterior processes ( Fig. 168 View FIGURES 163–169 )....................................................... 16
- Hypandrium without latero-posterior processes ( Fig. 71 View FIGURES 67–72 )..................................................... 17
16. Hypandrium with widely separated latero-posterior processes, directed outward ( Fig. 168 View FIGURES 163–169 ), epiproct with strongly convex anterior margin ( Fig. 166 View FIGURES 163–169 )............................................................... M. tukunare n. sp.
- Hypandrium with closely spaced latero-posterior processes, directed caudally, these are short, widened, with rounded lateralposterior margin and small tooth-shaped processes ( Fig. 107 View FIGURES 103–108 ); epiproct with concave anterior margin ( Fig. 106 View FIGURES 103–108 ).............................................................................................. M. chocoensis n. sp.
17. Phallosome very narrow basally ( Figs 72 View FIGURES 67–72 , 96 View FIGURES 91–96 , 173 View FIGURES 170–180 ).......................................................... 18
- Phallosome basally widened or greatly widened ( Figs 102 View FIGURES 97–102 , 150 View FIGURES 145–150 , 162 View FIGURES 157–162 , 173 View FIGURES 170–180 )....................................... 20
18. Hypandrium with small medial apical bifurcated process......................................... M. pennyi View in CoL (New)
- Hypandrium without medial apical bifurcated process ( Fig. 95 View FIGURES 91–96 )................................................ 19
19. Phallosome with external parameres apically briefly separated by membrane, endophallus elongated, flattened and apically more splayed ( Fig. 96 View FIGURES 91–96 )................................................................... M. caliensis n. sp.
- Phallosome with external parameres broadly fused medially and its endophallus is not elongated and apically more broadened ( Fig. 72 View FIGURES 67–72 )............................................................................ M. amacayacu n. sp.
20. Phallosome basally broadened, curved inward, apically rounded, with narrow posterior lobes ( Fig. 150 View FIGURES 145–150 ), endophallus membranous........................................................................................ 21
- Phallosome basally narrower than distally or membrane-bound side struts, posterior lobes broadened ( Figs 102 View FIGURES 97–102 , 156 View FIGURES 151–156 , 162 View FIGURES 157–162 )... .................................................................................................. 22
21. Epiproct with convex anterior margin, projecting onto the clunium; posterior process of paraprocts with rounded apex ( Fig. 148 View FIGURES 145–150 ); phallosome with widened side struts ( Fig. 150 View FIGURES 145–150 )............................................. M. katios n. sp.
- Epiproct semioval, with anterior margin straight to concave; paraprocts with posterior process acuminates; phallosome with narrow side struts ( Fig. 176 View FIGURES 170–180 )....................................................... M. minor View in CoL (New & Thornton)
22. Phallosome not piriform, side struts basally separate or widely united external parameres broad distally ( Fig. 162 View FIGURES 157–162 )....... 23
- Phallosome piriform, with side struts basally fused or joined by membrane ( Figs 102 View FIGURES 97–102 , 156 View FIGURES 151–156 ).......................... 24
23. Epiproct hexagonal, distal-lateral sides longer than others and heavily sclerotized; phallosome with side struts basally separated, joined by membrane, external parameres with internal margin gently curved and apically joined by membrane ( Fig. 174 View FIGURES 170–180 )....................................................................................... M. parvus Mockford View in CoL
- Epiproct semioval ( Fig. 160 View FIGURES 157–162 ); phallosome with side strut basally united by broad sclerotized area, external parameres with internal margin strongly curved, apically converging mesally ( Fig. 162 View FIGURES 157–162 )........................ M. putumayensis n. sp.
24. Phallosome with side struts basally fused ( Fig. 102 View FIGURES 97–102 ); forewing with length/width ratio of approximately 2.8 ( Fig. 97 View FIGURES 97–102 ).......................................................................................... M. caquetensis n. sp.
- Phallosome with side struts basally not fused, joined by membrane ( Fig. 156 View FIGURES 151–156 ); forewing with length/width ratio of approximately 2.4 ( Fig. 151 View FIGURES 151–156 )........................................................................... M. panchei n. sp.
25. Species in which only females are known................................................................. 26
- Species in which males and females or only males are known................................................. 29
26. Forewing with large depigmented area from the rear angle pterostigma-stigmasac, diagonal to the posterior margin, interrupted only by a point around the vein M before the intersection between the vein M and Cu1a (see Fig. 1 View FIGURES 1–5 in Williner 1910).......................................................................................... L. argentina Williner View in CoL
- Forewing without large depigmented area from the rear angle pterostigma-stigmasac, diagonal to the posterior margin.... 27
27. Vertex emarginated; forewing with M1 – M3 cells and apical half of R5 completely pigmented (see Fig. 175 View FIGURES 170–180 in New & Thornton 1975)....................................................................... L. pulchella View in CoL (New & Thornton)
- Vertex not emarginated, forewing with speckled pattern of coloration........................................... 28
28. Apical sclerite of subgenital plate with a medial group of setae and truncated apically................ L. timmei Mockford View in CoL
- Apical sclerite of subgenital plate without setae and rounded apically........................... L. thorntoni Mockford View in CoL
29. Phallosome with side struts fused at least at base, narrow or greatly enlarged, basally rounded; hypandrium variable in its shape ( Figs 11 View FIGURES 6–12 , 29 View FIGURES 25–30 , 41 View FIGURES 37–42 )..................................................................................... 32
- Phallosome with side strut separate, joined only by membrane ( Fig. 172 View FIGURES 170–180 ); hypandrium triangular, longer than wide....... 30
30. Hypandrium with lateral processes long and slender.............................................. L. varia (Navas) View in CoL
- Hypandrium with lateral processes short, pointed corners..................................................... 31
31. Hypandrium with lateral processes scarcely protruding beyond edges; forewing without clear band parallel to apical margin...................................................................................... L. onca Mockford View in CoL
- Hypandrium with lateral processes obviously protruding as pair points; forewing with clear band parallel to apical margin.................................................................................... L. conspersa Enderlein View in CoL
32. Hypandrium shorter than wide, bilobed apically ( Fig. 29 View FIGURES 25–30 ); phallosome widely enlarged, with rounded anterolateral angles, with medial sclerotization ( Fig. 30 View FIGURES 25–30 ), external parameres with elongated tooth-like processes on their inner margin................................................................................................... L. huilensis n. sp.
- Hypandrium as long as wide or longer than wide; phallosome elongated......................................... 33
33. Epiproct vase-shaped ( Fig. 179 View FIGURES 170–180 ), hypandrium triangular, phallosome with external parameres present as short, slender arms ( Fig. 170 View FIGURES 170–180 )...................................................................... L. punctipennis Mockford View in CoL
- Epiproct variable in shape, not vase-shaped; variable phallosome.............................................. 34
34. Epiproct with superficial dorsal flap ( Fig. 40 View FIGURES 37–42 ); hypandrium with extensive basally projecting sclerotization ( Fig. 41 View FIGURES 37–42 )...... 35
- Epiproct without superficial dorsal flap ( Figs 4 View FIGURES 1–5 , 10 View FIGURES 6–12 and 34 View FIGURES 31–36 ); hypandrium is different, not like the above ( Figs 11 View FIGURES 6–12 , 41 View FIGURES 37–42 )..... 37
35. Phallosome rounded basally ( Fig. 175 View FIGURES 170–180 )......................................................... L. ariasi View in CoL (New)
- Phallosome concave basally ( Fig. 42 View FIGURES 37–42 ).................................................................... 36
36. Phallosome broadened basally ( Fig. 42 View FIGURES 37–42 ), external parameres narrow, curved inward, sharply angled, aedeagus distally widened, endophallic membrane with well sclerotized apical spinous processes.............................. L. lienhardi n. sp.
- Phallosome narrow basally, external parameres distally widened, curved inward but not strongly angled as above, aedeagus apically bilobed, each lobe with elongated processes, endophallic membrane without apical spinous processes.................................................................................................... L. ariasi View in CoL (New)
37. Phallosome with side strut apically distinct, elongated, slender, or distally curved inward ( Fig. 12 View FIGURES 6–12 ); hypandrium U-shaped, usually longer than wide ( Fig. 11 View FIGURES 6–12 )....................................................... L. gorgonaensis n. sp.
- Phallosome with side strut apically undifferentiated in its apical part of the external parameres, hypandrium with variable shape.............................................................................................. 38
38. Phallosome sledgehammer-shaped, external parameres as two flattened and converging lobes ( Fig. 5 View FIGURES 1–5 )................. 39
- Phallosome with variable shape but not sledgehammer-shaped, variable external parameres ( Figs 24 View FIGURES 19–24 , 36 View FIGURES 31–36 and 48 View FIGURES 43–48 )......... 41
39. Phallosome with side strut basally close together ( Fig. 5 View FIGURES 1–5 )..................................... L. bonaverensis n. sp.
- Phallosome with side strut separate basally, bound together by a membrane ( Fig. 60 View FIGURES 55–60 ) or somewhat broadened and rounded basally ( Fig. 171 View FIGURES 170–180 ).................................................................................... 40
40. Phallosome with external parameres with small apical lobes, endophallus with large sclerotized area and membrane with abundant spicules as illustrated ( Fig. 60 View FIGURES 55–60 )..................................................... L. pearmani n. sp.
- Phallosome with external parameres without apical lobes as above, endophallus without sclerotized area and spicules as above ( Fig. 171 View FIGURES 170–180 )...................................................................... L. chelata Thornton & Woo View in CoL
41. Hypandrium with preapical constriction, forming a broad distal lobe ( Fig. 35 View FIGURES 31–36 ); phallosome with mesal endophallic process, more sclerotized and somewhat acuminate apically ( Fig. 36 View FIGURES 31–36 )...................................... L. iscuande n. sp.
- Hypandrium without preapical constriction ( Figs 23 View FIGURES 19–24 , 47 View FIGURES 43–48 ); phallosome without mesal endophallic process ( Fig. 48 View FIGURES 43–48 )....... 42
42. Phallosome with side strut widened, external parameres apically more broadened ( Fig. 48 View FIGURES 43–48 ), epiproct semioval ( Fig. 46 View FIGURES 43–48 )........................................................................................ L. occidentalis n. sp.
- Phallosome with side strut narrow, external parameres evenly broadened ( Fig. 24 View FIGURES 19–24 ); subtrapezoidal epiproct ( Fig. 22 View FIGURES 19–24 )............................................................................................ L. guapiensis n. sp.
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