Burmella, Godunko & Martynov & Staniczek, 2021

Godunko, Roman J., Martynov, Alexander V. & Staniczek, Arnold H., 2021, First fossil record of the mayfly family Vietnamellidae (Insecta, Ephemeroptera) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution, ZooKeys 1036, pp. 99-120 : 99

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scientific name


gen. nov.

Genus Burmella gen. nov. Figures 1 View Figure 1 , 2 View Figure 2 , 3 View Figure 3 , 4 View Figure 4 , 5 View Figure 5 , 6 View Figure 6 , 7 View Figure 7 , 8 View Figure 8 , 9 View Figure 9 , 10 View Figure 10 , Table 1

Type species.

Burmella paucivenosa sp. nov.

Derivation of name.

The generic name of female gender is a composition of " Burmar " as an ancient term for Myanmar, combined with " ella ", a common ending of generic names in mayflies, and especially so within Ephemerelloidea .


Adults of Burmella gen. nov. differ from other mayfly genera by the following combination of features: forewings (a) with small number of cross veins; (b) pterostigma with simple veins, not anastomosed; (c) CuP smoothly curved towards wing base; (d) two secondary bifurcate veins in cubital field; (e) at least several free marginal intercalaries along ventral margin; hind wings (f) strongly rounded, small, as long as 0.08-0.14 of forewing length; (g) small number of cross veins; (h) triad RS present or absent; no MA and MP triads; (i) no secondary branches of cubital veins; (j) costal process developed, rounded apically, situated centrally; abdomen (k) with vestigial gill sockets recognizable at least on segments II-VI; genitalia (l) with large median projection of styliger plate, widely rounded apically; (m) three distal segments of forceps strongly elongated and slender; segment II longest, 5 × as long as segment III; segments III and IV approximately of equal length; segment IV expanding apically; (n) penis lobes widely separated by V-shaped cleft; (o) no trace of paracercus. Additionally, in female (p) anterior part of eyes covered by anterolaterally expanded clypeal shield.

Subimago and larva unknown.

Species composition.

Burmella paucivenosa sp. nov. (SMNS; BU-179); Burmella clypeata sp. nov. (SMNS; BU-321).

Locality and horizon.

Hukawng Valley, Kachin State, Myanmar (Burma); Cenomanian, mid-Cretaceous.