Chassalia macrocarpa T.Y.Yu, 2021

Turner, Ian M. & Cheek, Martin, 2021, Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species, European Journal of Taxonomy 738, pp. 1-60 : 40

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scientific name

Chassalia macrocarpa T.Y.Yu

sp. nov.

Chassalia macrocarpa T.Y.Yu sp. nov.

Figs 22–23 View Fig View Fig


Differs from all other Bornean species of Chassalia in having relatively large fruits (10–12 × 10 × 5 mm); differs from other species of Chassalia from Mt Matang in having larger rachis and capitulum bracts, rachis bracts usually round or acuminate with long acumen which covers the flower buds, and in fruits becoming pale yellowish brown when dry.


The epithet was chosen because, within Borneo, this species of Chassalia has large fruits (1.2 × 1 × 0.5 cm).


MALAYSIA – Borneo, Sarawak • 1 st Division, Kuching, Matang , Path to Bukit Tanduk ; alt. 750 m; 23 Apr. 1987; Bernard Lee Meng Hock S 54027 View Materials ; holotype: K [ K001129715 ]!; isotype: SAN .

Additional material

MALAYSIA – Borneo, Sarawak • 1 st Division, Kuching, Matang , by trail; alt. 450 m; 27 Oct. 1929; M. & J. Clemens 22373; K!, SAR 28 Oct. 1929; M. & J. Clemens 22373; K!, SAR alt. 500 m; 31 Jul. 1963; Paul Chai S 18534 View Materials ; A, K!, L, SAN, SAR Feb. 1915; H. N. Ridley s.n.; K !.


Shrub, 1–1.5 m tall, glabrous. Internodes (1–) 1.5–4.5 cm × 3–4 mm, lower internodes terete, upper internode usually drying flattened, canaliculate, broadening at apex. Leaf-blades narrowly elliptic to narrowly lanceolate, 15–24 × 2–5.5 cm, margin entire, apex acute to acuminate, base cuneate; midrib slightly raised above, domatia absent; secondary nerves 12–16 on each side of the midrib, prominent above; tertiary nerves not visible. Petiole very short, canaliculate, usually slightly winged, 10–25 × 1–2 mm. Stipule not sheathing, broadly flabellate, 1–2 × 1–1.5 cm, apex usually broken. Inflorescence a compound cyme, (1–) 1.8–5 cm long; peduncle (0.5–) 1.5–3.5 cm long; Bracts of second internode of rachis and second branches 2, round to elliptic, 1.2 × 1 cm, usually slightly sheathing at base, apex obtuse, sometimes acuminate with acumen 5 mm long, second branches usually very short to absent, bracts of capitulum elliptic, 10–12 × 5–10 mm. Open flowers not seen. Fruits purplish blue when young, blue when ripe, light yellowish brown when dry, obovoid, 10–12 × 10 × 5 mm. Pyrenes plano-convex, dorsal surface convex, with two adjacent parallel longitudinal crests, each 0.3 mm wide, curved, 0.3 mm apart along their length, forming a longitudinal groove from apex to base between the two crests; ventral surface concave, centre slightly elevated, margin with an endocarpal outgrowth 0.5–0.8 mm wide and a groove between the edge of endocarp and endocarpal outgrowth 0.3–0.5 mm wide. Raphal opening at base of the endocarpal outgrowth, round, 0.2–0.3 mm diameter, 0.2–0.3 mm from the edge of the pyrene wall, apex with a bony spine, flabellate, erect, 1.5 × 0.3 × 0.6–0.7 mm. PGS not detected. Seed not seen.

Distribution, habitat

Borneo: Malaysia, Sarawak, 1 st Division, Kuching, Matang. Mixed dipterocarp forest, alt. 450– 750 m.


Royal Botanic Gardens


Forest Research Centre


Botanische Staatssammlung München


University of the Witwatersrand


Department of Forestry


Harvard University - Arnold Arboretum


Nationaal Herbarium Nederland, Leiden University branch


University of Helsinki


Nanjing University