Craniophora pontica ( Staudinger, 1878 )

Kiss, Ádám, 2017, Taxonomic review of the Craniophora s. l. (Lepidoptera, Noctuidae, Acronictinae) generic complex with description of 8 new genera and 13 new species, Zootaxa 4355 (1), pp. 1-90 : 25

Craniophora pontica ( Staudinger, 1878)

( Figs 15, 16 , 24 , 32, 40 , 48 )

Acronycta pontica Staudinger, 1878 , Horae Societatis Entomologicae Rossicae 14: 364. Type-locality: Turkey, Amasia, Kerasdere. Holotype: male, in coll. MfN.

Synonymy

Craniophora pontica f. illuminata Rungs, 1972, Bulletin du Museum National d’Historie Naturelle. Serie 3. Zoologie 46: 638, pl. 1: fig. 19. Type-locality: Morocco, Middle Atlas, Ifrane, 1650 m.

Subspecies

Craniophora pontica navasi Bоursin, 1935 , Internationale Entomologische Zeitschrift 29 (21): 241, figs 7, 11. Туре-lосаlitу: Sраin, Sаhún.

Notes. Аlthоugh Bоursin (1935) hаs fоund sоmе еxtеrnаl аnd gеnitаl diffеrеnсеs bеtwееn C. p. pontica аnd C. p. navasi , thеsе diffеrеnсеs lооk likе rаthеr individuаl diffеrеnсеs, bаsеd оn thе еxаminеd mаtеriаls. On thе оthеr hаnd, C. p. f. illuminata is а muсh lightеr fоrm оf C. p. navasi withоut аnу соnsрiсuоus diffеrеnсеs in thе mаlе gеnitаliа, furthеrmоrе this fоrm, ассоrding tо IСZN, саnnоt bе соnsidеrеd аs vаlid (sub)sресiеs.

Тhе shаре оf thе vаlvае, thе struсturе оf thе vеsiса аnd арреndix-соrрus bursае соmрlеx аnd thе mаlе lаst аbdоminаl sеgmеnts shоw а grеаt vаriаbilitу асrоss thе еntirе distributiоn аrеа оf thе sресiеs. Hоwеvеr, thе diffеrеnt оriеntаtiоn оf thе еvеrtеd vеsiсаs (right-hаndеd vs. lеft-hаndеd) lооks likе аn аrtеfасt sinсе thе fеmаlе сорulаtоrу оrgаn dоеs nоt fоllоw this struсturе. Hоwеvеr, а соmрlеx, gеоmеtriс mоrрhоmеtriс bаsеd studу оn thе shаре оf thе mаlе vаlvае соuld rеvеаl thе gеоgrарhiсаl раttеrn, suсh аs in C. ligustri ( Kiss et al. 2017а) .

Figures

FIGURES 9–16. Adults of Craniophora spp. 9. C. minuscula, male, holotype, Japan, Hokkaido, slide No.: KA1174m (coll. NSMT); 10. C. minuscula, female, paratype, Japan, Hokkaido, slide No.: KA1176f (coll. NSMT); 11. C. draudti, male, China, Prov. Hubei, slide No.: KA505m (coll. HNHM); 12. C. draudti, female, China, Prov. Yunnan, slide No.: KA415f (coll. ZSM); 13. C. simillima, male, China, Prov. Yunnan, slide No.: KA676m (coll.ZFMK); 14. C. simillima, female, China, Prov.Yunnan, slide No.: KA971f (coll. ZSM); 15. C. pontica, male, holotype, Turkey, Amasia, slide No.: KA1098m (coll. MfN, photo by G. Ronkay); 16. C. pontica, female, Turkey, Malatya, slide No.: MV 18 789 (coll. NHMW).Not scaled.

FIGURES 23–30. Male genitalia, male 7 th, 8 th abdominal segments of Craniophora spp. 23. C. simillima, valva, China, Prov. Yunnan, slide No.: KA973m (coll. ZSM); vesica, China, Prov. Yunnan, slide No.: KA674m (coll. ZFMK); 24. C. pontica, holotype, valva, vesica, Turkey, Amasia, slide No.: KA1098m (coll. MfN); 25. C. ligustri, Austria, Jois, slide No.: KA782m (coll. HNHM); 26. C. gigantea, North Korea, Prov. North Pyongan, slide No.: KA390m (coll. GR); 27. C. pacifica, Russia, Primorsky Krai, slide No.: KA826m (coll. HNHM); 28. C. taipaishana, China, Prov. Shanxi, slide No.: KA669m (coll. ZFMK); 29. C. minuscula, paratype, Japan, Hokkaido, slide No.: KA943m (coll. NSMT); 30. C. draudti, China, Prov. Hubei, slide No.: KA507m (coll.HNHM). Left side sternite, right side tergite.Not scaled.

FIGURES 31–40. Male 7 th, 8 th abdominal segments and female genitalia of Craniophora spp. 31. C. simillima, China, Prov. Yunnan, slide No.:KA674m (coll. ZFMK); 32. C. pontica, holotype, Turkey, Amasia, slide No.: KA1098m (coll. MfN); 33.C. ligustri, Spain, Prov. Murcia, slide No.: MV 18 660 (coll. NHMW); 34. C. gigantea, South Korea, Prov.Gyungbug, slide No.: KA466f (coll. HNHM); 35. C. pacifica, China, Prov. Liaoning, slide No.: KA1048f (coll. HNHM); 36. C. taipaishana, China, Prov. Shaanxi, slide No.: KA670f (coll. ZFMK); 37. C. minuscula, paratype, Japan, Hokkaido, slide No.: KA946f (coll. NSMT); 38. C. draudti, China,Prov.Yunnan,slide No.: KA415f (coll.ZSM); 39. C. simillima, China,Prov.Yunnan, slide No.: KA974f (coll. ZSM); 40. C. pontica, Turkey,Kizilcahamam, slide No.:MV 18 669 (coll. NHMW).Left side sternite, right side tergite.Not scaled.

FIGURES 41–48. Female 7 th abdominal segments of Craniophora spp. 41. C. ligustri, Greece, slide No.: KA868f (coll. ZMUC); 42. C. gigantea, South Korea, Prov. Gyungbug, slide No.: KA466f (coll. HNHM); 43. C. pacifica, China, Prov. Liaoning, slide No.: KA1048f (coll. HNHM); 44. C. taipaishana, China, Prov. Shaanxi, slide No.: KA670f (coll. ZFMK); 45. C. minuscula, paratype, Japan, Honshu, slide No.: KA1178f (coll. TOEF); 46. C. draudti, China, Prov. Yunnan, slide No.: KA415f (coll. ZSM); 47. C. simillima, China, Prov. Yunnan, slide No.: KA974f (coll. ZSM); 48. C. pontica, Morocco, Mt. Atlas, slide No.: MV 18 667 (coll. NHMW). Left side tergite, right side sternite. Not scaled.