Messatoporus mesonotator Kasparyan & Ruíz 2005, 2008

Santos, Bernardo F. & Aguiar, Alexandre P., 2013, <strong> Phylogeny and revision of <em> Messatoporus </ em> Cushman (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of sixty five new species </ strong>, Zootaxa 3634 (1), pp. 1-284 : 151-152

publication ID	https://doi.org/ 10.11646/zootaxa.3634.1.1
publication LSID	lsid:zoobank.org:pub:81906FA2-FB3C-4F02-9AF9-449BC73F3BEE
persistent identifier	https://treatment.plazi.org/id/3629866A-FF08-2ED0-B8BF-FF68FC76B66E
treatment provided by	Felipe
scientific name	Messatoporus mesonotator Kasparyan & Ruíz 2005
status

Treatment

Messatoporus mesonotator Kasparyan & Ruíz 2005 View in CoL

( Figs 78 View FIGURES 76–80 , 197 View FIGURES 189–197 , 236 View FIGURES 236–241 , 344 View FIGURES 337–344 )

* Messatoporus mesonotator Kasparyan & Ruíz 2005: 107 View in CoL , 116. Description, distribution, keyed. Holotype ♀ (UATM, not examined). Type data: México, Tamaulipas, Gómes Farías, Alta Cima, 900 m, Malaise trap, 7–14.X.2000, S. Hernández A. Messatoporus mesonotator: Kasparyan & Ruíz 2008: 309 View in CoL , 312, 329–330, 373. Description, distribution, figure, keyed.

Female. Fore wing 7.5 mm. HEAD: Mandible with moderately sparse and long hairs; MLW 1.74; MWW 0.31; dorsal tooth in front view projected upwards, long, ventral tooth indistinct. MSM 0.33. Clypeus slightly and more or less uniformly convex, centrally straight; CHW 2.53; CWW 2.24; apical area medially straight, laterally with distinct triangular lobes, its margin regular, not raised. Antenna with 26 flagellomeres; flagellum uniform; apex of apical flagellomere much narrower than base, distinctly tapered. Supra-antennal area shiny, scarcely punctate, distinctly striate near antennal sockets, ventrally distinctly concave, without median line. Occipital carina uniformly curved, apically very faint, meeting hypostomal carina far from mandible base. Temple and gena moderately wide, gena somewhat swollen and projected (as in Fig. 261 View FIGURES 260–271. 260–261 ).

THORAX: Pronotum glabrous, mat, scarcely punctate, ventrally moderately striate; epomia weak, distinct only after diverging from pronotal collar, short, ending far from dorsal margin of pronotum, after diverging from pronotal collar distinctly curved. Mesoscutum moderately and uniformly convex, ovoid, 1.19 × as long as wide, densely covered with short hairs, shiny, densely punctate; notaulus reaching about 0.68 of mesoscutum length, moderately impressed, surface over notaulus with weak and short transverse wrinkles; scuto-scutellar groove moderately deep, with vestigial longitudinal wrinkles. Subalar ridge weakly projected; epicnemial carina reaching about 0.7 of distance to subalar ridge, irregular; sternaulus moderately strong, wider on posterior apex, strongly sinuous, with weak vertical wrinkles; scrobe moderately deep; mesopleural suture with weak longitudinal wrinkles along entire length. Mesosternum medially with short transverse wrinkles; median portion of posterior transverse carina of the mesothoracic venter short, slightly arched forwards. Transverse sulcus at base of propodeum very wide, about 0.84 × as long as anterior area of propodeum, laterally with weak longitudinal wrinkles, medially not wrinkled; metapleuron impunctate, covered with the same wrinkling of propodeum, however less regular, covered with moderately dense, very short hairs; juxtacoxal carina represented by very short ridges. Fore tibia weakly swollen. Mesal lobe of t4 with a distinct cluster of stout bristles. Hind coxa punctate-coriarious.

PROPODEUM: 1.15 × as long as wide, shiny, sparsely pilose; anterior area scarcely punctate; spiracle elongate, SWL 4.44; anterior transverse carina medially slightly arched forwards. Propodeal wrinkles moderately strong, closely spaced, anteriorly complete, posteriorly mostly incomplete and overlapped, distinctly curved backwards; posterior transverse carina distinct as very weak sublateral crests.

WINGS: Fore wing vein 1-Rs+M distinctly sinuous, with bulla placed on basal 0.25; crossvein 1m-cu more or less uniformly curved, limit between 1m-cu and 1-Rs+M more or less traceable; vein 1M+Rs anteriorly straight, posteriorly weakly curved; fore wing crossvein 1 cu-a basad of 1M+Rs by 0.4 of its own length; vein 2Cua 0.57 × as long as crossvein 2cu-a; bulla of crossvein 2m-cu placed mostly on posterior half; cell 1+2Rs small, APH 0.55, pentagonal, about as high as wide, AWH 0.97; crossvein 3r-m fully spectral, 2r-m and 3r-m subparallel, about same length; vein 3-M distinctly shorter than 2-M; 4-Rs very weakly sinuous; 4-M semi-spectral, straight. Hind wing vein 1-M forming distinctly obtuse angle with vein Cua; vein 2-Rs entirely tubular, apically weaker, reaching wing margin; HW1C 0.93; vein Cub distinctly convex on posterior half, forming straight angle with Cua; vein 2-1A reaching 0.89 of distance to posterior wing margin.

METASOMA: First tergite moderately long, about 0.37 × as long as T2–8, somewhat depressed, ventrolaterally angled, dorsally sparsely pilose, laterally moderately pilose; T1LW 3.75; T1WW 1.83; spiracle on 0.48 of its length, distinctly prominent; median depression elongate, very shallow; lateral and median posterior depressions absent; ventrolateral carina absent, but traceable as limit between lateral and ventral portions. T2LW 1.09; T2WW 2.26; thyridium longer than wide, without small circular depression just behind it; T2–8 smooth between punctures, densely and uniformly pilose. OST 0.76; ovipositor moderately stout, straight, basally cylindric, apically distinctly depressed; dorsal valve with four weak ridges; ventral valve apex with 11 teeth, apical teeth progressively more closely spaced; surface anterior to first tooth rugulose.

Color. Head, mesosoma and metasoma black and whitish. Head: black; clypeus, basal spot at mandible, mouthparts, malar space, supra-clypeal area, scape ventrally and complete orbital band, whitish (235,220,182); f8–14 entirely white, f7 and f15 mostly white. Mesosoma: black; collar and dorsal margin of pronotum connected, propleuron, U-shaped mark at mesoscutum, scutellar carina, scutellum, axillar carinae, postscutellum, tegula, subalar ridge, mesopleuron except borders and scrobe, mesosternum, dorsal division of metapleuron, metapleural triangle, metapleuron except posterior apex and arc-shaped, quadratic mark on propodeum, yellowish (222,199,119); fore and mid coxae and first trochanters whitish, coxae with basal and apical brown marks, first trochanters dorsally with basal brown mark; fore femur, tibia and tarsus whitish, femur with weak dorsal brownish stripe, tibia with ventral brownish stripe, t5 blackish; mid femur ventrally whitish, dorsally pale orange; mid tibia and tarsus whitish, t5 blackish; hind coxa whitish with three longitudinal stripes, anterior and posterior ones reaching 0.5 of its length, lateral one reaching coxa apex; hind trochanters pale orange (198,104,042) with basal brownish marks; hind femur pale orange; basal 0.8 of hind tibia pale yellow, apical 0.2 dark brown; basal 0.25 of t1 dark brown, remainder of hind tarsus white. Metasoma: black; anterior 0.5 dorsally and posterior 0.2 of T1 whitish; S1 whitish; T2–8 and S2–6 with posterior whitish stripes.

Male. Fore wing 5.57–7.00 mm long. Very similar to female, except for the following: epicnemium ventrally whitish; blackish marks at tibia slightly more extensive.

Variation. Fore wing 6.50–8.78 mm long. Mandible sometimes mostly whitish; antenna sometimes with 25 flagellomeres; pronotum sometimes almost completely, or at males, completely, yellow; anterior portion of hind coxa sometimes pale orange; hind femur dorsally sometimes with small brownish basal marks; T2 often with anterior whitish spot; black areas at S5–6 sometimes reduced to small lateral brownish marks.

Comments. Very similar to M. jalapa , from which it can be differentiated by the yellow mark on mesopleuron connected to yellow subalar ridge (vs. not connected); T1 laterally black at anterior 0. 8 (vs. laterally entirely yellow); ovipositor moderately stout, dorsal valve with four weak ridges (vs. slender, dorsal valve without ridges); ventral tooth of mandible indistinct (vs. distinct and triangular); and supra-antennal area distinctly striate near antennal sockets (vs. not striate). Also somewhat similar to M. maculipes , from which it can be differentiated mainly by the yellow mark at mesoscutum U-shaped (vs. subcircular); anterior area of propodeum entirely black (vs. with lateral yellow marks); T1 laterally black at anterior 0.8 (vs. laterally entirely yellow); sternaulus posterior transverse carina of propodeum distinct as weak sublateral crests (vs. entirely absent); and ovipositor dorsal valve with four weak ridges (vs. five moderately strong ridges).

Biology. Unknown.

Material examined. 7 ♀, 6 ♂♂. 1 ♂ from MEXICO, Chiapas, Mpio, San Cristobal, San Felipe , 2195 m, 20–25.VII.1990 , Malaise trap, RWJones ( TAMU) . 1 ♀ from COSTA RICA, Guanacaste, San Rosa Park, Dry hill, 7.VII.1978 , DHJanzen; 1 ♂, same data except 11.XI.1977 ; 1 ♂, same data except 10.VII.1977 ; 1 ♂, same data except 4.IX.1977 ; 1 ♂, same data except 15.IX.1977 ; 1 ♂, same data except 26.VIII.1977 . 1 ♀ from COSTA RICA, Puntarenas, Monteverde , 5.I.1962 , CPalmer; 1 ♀, same data except 17–19.II.1968 ; 1 ♀ from COSTA RICA, San José, Escazú , 22.V.1987 , Townes & Townes ( AEIC) . 1 ♀ from PANAMA, Canal Zone, Barro Colorado Island , IV–V.1942 , JZetek; 1 ♀, same data except IX–X.1939 ; 1 ♀, same data except X.1939 ( USNM).