Messatoporus depressifrons Santos, 2013
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publication ID |
https://doi.org/ 10.11646/zootaxa.3634.1.1 |
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publication LSID |
lsid:zoobank.org:pub:81906FA2-FB3C-4F02-9AF9-449BC73F3BEE |
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persistent identifier |
https://treatment.plazi.org/id/3629866A-FFF9-2E2E-B8BF-FD93FB18B205 |
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treatment provided by |
Felipe |
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scientific name |
Messatoporus depressifrons Santos |
| status |
sp. nov. |
Messatoporus depressifrons Santos , sp. nov.
( Figs 73 View FIGURES 71–75 , 237 View FIGURES 236–241 , 274 View FIGURES 272–278. 272–273 , 318 View FIGURES 313–320 )
Female. Fore wing 8.91 mm. HEAD: Mandible densely covered with long hairs; MLW 2.29; MWW 0.42; dorsal tooth in front view projected upwards, much longer than ventral tooth; ventral tooth triangular. MSM 0.25. Clypeus slightly and more or less uniformly convex, centrally concave; CHW 2.56; CWW 3.95; apical area medially concave, laterally slightly projected, its margin very sharp, raised. Antenna with 27 flagellomeres; flagellum strongly flattened and compressed on apical 0.3; apex of apical flagellomere about as wide as base, not distinctly tapered. Supra-antennal area shiny, with moderately dense punctation, not striate near antennal sockets, ventrally and around ocelli distinctly concave, medially with a distinct depression but without a distinct line. Occipital carina uniformly curved, fading out at distance about as long as basal width of mandible from the hypostomal carina. Temple and gena moderately wide, gena somewhat swollen and projected (as in Fig. 261 View FIGURES 260–271. 260–261 ).
THORAX: Pronotum glabrous, shiny, impunctate, ventrally with strong wrinkles; epomia moderately strong, distinct only after diverging from pronotal collar, short, ending far from dorsal margin of pronotum, after diverging from pronotal collar distinctly curved. Mesoscutum moderately convex, flattened posteriorly, ovoid, 1.37 × as long as wide, sparsely pilose, shiny, minutely punctulate; notaulus reaching about 0.76 of mesoscutum length, deeply impressed, surface over notaulus not wrinkled; scuto-scutellar groove deep, with distinct longitudinal wrinkles. Subalar ridge weakly projected; epicnemial carina reaching about 0.7 of distance to subalar ridge, irregular; sternaulus moderately strong, wider on posterior apex, strongly sinuous, smooth; scrobe moderately deep; mesopleural suture ventrally with strong longitudinal wrinkles, wrinkles dorsally weak. Mesosternum medially with short transverse wrinkles; median portion of posterior transverse carina of the mesothoracic venter short, arched backwards. Transverse sulcus at base of propodeum moderately wide, about 0.59 × as long as anterior area of propodeum, impunctate; metapleuron moderately punctate, without transverse wrinkles, moderately pilose; juxtacoxal carina absent. Fore tibia distinctly swollen. Mesal lobe of t4 with a distinct cluster of stout bristles. Hind coxa with weak punctures separated by more than their own diameter.
PROPODEUM: 1.04 × as long as wide, shiny, moderately pilose; anterior area scarcely punctate; spiracle elongate, SWL 6.25; anterior transverse carina medially straight. Propodeal wrinkles strong, closely spaced, anteriorly faint at median portion, posteriorly partly complete and partly incomplete, mostly slightly curved or somewhat sinuous; posterior transverse carina entirely absent.
WINGS: Fore wing vein 1-Rs+M distinctly sinuous, with bulla placed on basal 0.4; crossvein 1m-cu irregular, limit between 1m-cu and 1-Rs+M more or less traceable; vein 1M+Rs anteriorly straight, posteriorly weakly curved; fore wing crossvein 1 cu-a basad of 1M+Rs by 0.49 of its own length; vein 2Cua 0.32 × as long as crossvein 2cu-a; bulla of crossvein 2m-cu placed mostly on posterior half; cell 1+2Rs small, APH 0.71, pentagonal, about as high as wide, AWH 0.94; crossvein 3r-m fully spectral, 2r-m and 3r-m subparallel, about same length; vein 3-M about as long as 2-M; 4-Rs somewhat irregular, apically slightly upcurved; 4-M tubular, irregular. Hind wing vein 1-M forming distinctly obtuse angle with vein Cua; vein 2-Rs tubular, apically spectral, reaching wing margin; HW1C 1.03; vein Cub distinctly convex on posterior half, forming straight angle with Cua; vein 2-1A reaching 0.81 of distance to posterior wing margin.
METASOMA: First tergite moderately long, about 0.45 × as long as T2–8, strongly depressed, ventrolaterally rounded, dorsally glabrate, laterally moderately pilose; T1LW 3.3; T1WW 1.74; spiracle on 0.55 of its length, distinctly prominent; median depression elongate, very shallow; lateral and median posterior depressions absent; ventrolateral carina absent, but traceable as limit between lateral and ventral portions. T2LW 1.29; T2WW 1.8; thyridium longer than wide, with small circular depression just behind it; T2–8 covered with small, shallow foveae, densely and uniformly pilose. OST 1.24; ovipositor moderately stout, straight, basally cylindric, apically slightly depressed; dorsal valve with four moderately strong ridges; ventral valve apex with 8 teeth, apical teeth progressively more closely spaced; surface anterior to first tooth rugulose.
Color. Head, mesosoma and metasoma black and whitish. Head: black; clypeus, basal 0.4 of mandible, mouthparts, supra-clypeal area, ventral spot at scape and complete orbital band, light yellow (226,208,098); f5–8 entirely white, f4 partially white, f9 almost entirely white and f10–14 with dorsal white longitudinal band. Mesosoma: black; very narrow mark at collar, posterior 0.5 of dorsal margin of pronotum, propleuron, subcircular spot on mesoscutum, scutellar carina, scutellum, axillar carinae, postscutellum, tegula, subalar ridge, central mark on mesopleuron, mesosternum, dorsal division of metapleuron, metapleuron except posterior apex and arc-shaped, quadratic mark on propodeum, light yellow; fore and mid coxae and first trochanters whitish, coxae with apical dark brown marks, mid one also with posterior face dark brown; first trochanters with basal dark brown mark; fore and mid second trochanter, femur, tibia and tarsus bright yellow (222,153,049), with strong dorsal brown stripes on femora, fore one with weak ventral stripe; tibiae with ventral brown marks, fore one on basal 0.8, mid one on basal 0.3; mid tarsus fuscous, fore and mid t5 blackish; hind coxa whitish with three longitudinal stripes, anterior one reaching 0.5 of its length, lateral and posterior ones reaching coxa apex; hind first trochanter whitish with basal blackish mark; second trochanter blackish, with apical 0.2 dark yellow; hind femur dorsal and anterior faces bright yellow, ventral face blackish, posterior face blackish on basal 0.1 and apical 0.2, medially bright yellow; hind tibia bright yellow, hind tarsus basally bright yellow, whitish towards apex. Metasoma: black; T1 laterally with anterior and posterior 0.2 whitish, dorsally anterior 0.5 whitish; T2–7 with posterior and lateral whitish stripes, T8 only with lateral stripes; S1–6 whitish, S2–5 with progressively smaller brown lateral marks.
Male. Unknown.
Variation. Fore wing 6.94–9.74 mm long. Arc-shaped mark on propodeum anteriorly either rounded or somewhat quadratic; hind tibia sometimes with very small blackish apical mark. Specimen from La Rioja with anterior propodeal wrinkles weak and median depression on T1 very strong.
Comments. Messatoporus depressifrons is part of a group of four very close species, which apparently form a monophyletic group ( Figs 71–75 View FIGURES 71–75 ; see Comments under M. compressicornis ). Among these species, M. depressifrons can be differentiated from M. keraiopetalus sp. nov. by having clypeus very slightly and more or less uniformly convex (vs. basally strongly convex, emarginated in its mid-length, abruptly flattened); supra-antennal area with a median depression (vs. with median smoother area); epomia moderately strong, distinct only after diverging from collar (vs. weak but complete); transverse sulcus of propodeum moderately long, 0.59 × as long as anterior area (vs. very long, 1.07 × as long as anterior area of propodeum); and fore wing vein 3-M and 2-M with about the same size (vs. 3-M distinctly longer).
It can be isolated from M. compressicornis by the hind femur ventrally black, dorsally deep yellow (vs. entirely orange); fore and mid femora dorsally with strong longitudinal brown stripes (vs. entirely orange); ovipositor dorsal valve with four moderately strong ridges (vs. five); supra-antennal area with median carina indistinct, represented only by a smoother area (vs. with slight median carina); and mesosternum medially with short transverse wrinkles (vs. without transverse wrinkles).
It can be separated from M. nigriscapus sp. nov. by the orbital band complete (vs. shortly interrupted); mandible basally yellow (vs. entirely black); collar yellow (vs. black); supra-antennal area with median depression (vs. with slight median carina); median portion of posterior transverse carina of the mesothoracic venter arched backwards (vs. arched forwards); and spiracle of propodeum elongate, SWL 4.86 (vs. elliptic, SWL 3.37)
Biology. One specimen from Moca, Guatemala, was reared from a nest of Pachymenes obscurus Smith ( Vespidae , Eumeninae).
Etymology. From the Latin depressus, meaning “depressed”, and frons; in reference to the longitudinal depression at supra-antennal area.
Material examined. 4 ♀. Holotype ♀ from GUATEMALA, Peten, Santa Elena , 120–160 m, VIII.1976, NLHKrauss ( AMNH). Pinned. Apical third of right antenna missing; otherwise in good shape.
Paratypes: 1 ♀ from GUATEMALA, Guatalon , Moca, III–IV.1931, “SC/ from nest of Pachymenes obscurus .”
1 ♀ from EL SALVADOR, San Salvador, 30.IV.1957, Pab, N. 714-250 ( USNM) . 1 ♀ from COSTA RICA ,
Puntarenas, La Rioja, 26.V.1956, LCKuitert, traps ( FSCA). Distribution. Recorded from three localities in Guatemala and Costa Rica ( Fig. 318 View FIGURES 313–320 ) .
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