Paramblynotus alexandriensis Buffington & van Noort, 2013

Noort, Simon van & Buffington, Matthew L., 2013, Revision of the Afrotropical Mayrellinae (Cynipoidea, Liopteridae), with the first record of Paramblynotus from Madagascar, Journal of Hymenoptera Research 31, pp. 1-64 : 28-30

Paramblynotus alexandriensis Buffington & van Noort sp. n. Figures 13

Type material.

HOLOTYPE. Female: South Africa, [first label] Alexandria, Cape Province, 22.2.1962, ACC. PE 857; [second label] with Curculionid in log of Ptaeroxylon obliquum ; [third label] (Hym. Cynipoidea , Mesocynipinae ) Paramblynotus Cameron, 1908, sp., det. Michael Soderlund, 1994; [fourth label] red holotype label (SANC). PARATYPES. 3F: Same data as holotype. Deposited in SANC, SAMC, and USNM.

Distribution.

South Africa.

Etymology.

Named after the Alexandria Forest, which now forms part of the Greater Addo Elephant National Park.

Diagnosis.

Belongs to the Paramblynotus trisetosus clade within the Paramblynotus trisetosus species-group of Liu et al. (2007). Female with 13 segmented antennae; male unknown. Ocellar plate present, mound-like; occiput concave in dorsal view. Mesoscutum deeply foveate, notaulices complete ( Fig. 13D ); upper mesopleuron entirely smooth, glabrous ( Fig. 13C ). T6 largest, T8 slightly exposed ( Fig. 13F ). Wings clear, no banding present ( Fig. 13A ). Most similar to Paramblynotus fuscapiculus , but distinguished by: the coloration of the female flagellum (terminal segment dark in Paramblynotus alexandriensis ( Fig. 13A ); terminal two segments dark in Paramblynotus fuscapiculus ); and setation of T9: in Paramblynotus alexandriensis , a dense brush of setae is present ( Fig. 13F ); in Paramblynotus fuscapiculus , T9 is glabrous or with only very short, appressed setae.

Description.

FEMALE. Length 2-2.5 mm. Head, mesosoma and metasoma dark brown; antennae and legs light yellow; terminal segment of antennae dark brown ( Fig. 13A ). Wings transparent ( Fig. 13A ). Entire head with the exception of the genae and occiput strongly pubescent ( Fig. 13E ). Eyes prominent, bulbous, but not laterally extended much beyond outer margin of genae in anterior view ( Fig. 13E ). Antenna 13 segmented; F1 shorter than F2; flagellum not widening toward apex. Vertex foveate, distinct carinae absent; ocellar plate not raised; ocelli large, their diameter as great as distance between lateral and median ocellus ( Fig. 13D ). Face punctate-rugose, humped between toruli and clypeal margin, protruding in lateral view; antennal scrobe mostly smooth with minute punctuation. Weakly keeled medial carina present between toruli extending towards median ocellus ( Fig. 13E ). Occiput concave in dorsal view, smooth without a carina. Lower face with strong excavations (with weak vertical carinae) defining upper clypeal margin, and containing anterior tentorial pits ( Fig. 13E ). Clypeus gently strigate. Genae with distinct foveae along eye margin, punctate-rugose and densely pubescent between these foveae and genal carina ( Fig. 13C ). Mesosoma strongly pubescent. Single submedian pronotal depression present. Anterior plate of pronotum puberulous. Pronotum dorsomedially with swollen rim without any crest. Lateral carinae of pronotum strong, fading dorsomedially. Lateral surface of pronotum foveate ( Fig. 13C ). Dorsal pronotal area smooth with minute punctures. Mesoscutum deeply foveate, setose; notaulices complete, extending to anterior margin of mesocutum; median mesoscutal impression absent ( Fig. 13D ). The two scutellar foveae each with three subcarina creating a transverse row of 8 longitudinally elongate subfoveae. Scutellum medially foveate, sparsely setose, peripherally areolate-punctate ( Fig. 13D ). Posterior mesoscutum and scutellum contiguously rounded in lateral view. Mesopleural triangle defined without ventral curved carina, strongly pubescent; upper mesopleuron glabrous, smooth, anteriorly and ventrally pubescent with minute punctures; median longitudinal impression percurrent with evenly spaced transverse carinae; speculum glabrous, smooth ( Fig. 13C ).

Metanotal-propodeal complex strongly excavated, excavations bordered by strong carinae. Metepisternum dorsally excavated with pubescence, medially polished with indications of minor rugose remnants, ventrally pubescent. Dorsellum with two strong medial foveae; laterally strongly excavated with fine pubescence in lateral depressions. Lateral propodeal carina present; median longitudinal propodeal carina well-defined, crossed by wrinkled transverse and longitudinal carinae extending onto nucha. Rs+M of forewing defined, but nebulous where it arises from basal vein at posterior third ( Fig. 13A ). Marginal cell 2.5 times as long as wide. Abdominal petiole short, longitudinally striate, 0.5 times as long as wide in dorsal view ( Fig. 13C ). Posterior ventral margin of metasomal T7 gently sinuate ( Fig. 13F ). T8 well exposed, with a patch of scattered long setae posteriorly ( Fig. 13F ). Ventral portions of T2-T7 covered by sternum 3. Tergites dorsally finely punctate; laterally and ventrally polished. All legs sparsely punctuate, strongly pubescent, except metacoxae dorsally glabrous, smooth ( Fig. 13F ). Mesotibial outer spur shorter than inner spur; metatibial spurs subequal in length. Ratio of first metatarsomere to the remaining 4 metatarsomeres combined: 0.67.

MALE. Unknown.

Figures

Figure 13. Paramblynotus alexandriensis sp. n., holotype female. A lateral habitus B dorsal habitus C head and mesosoma, lateral view D head and mesosoma, dorsal view E head, anterior view F metasoma, lateral view.