Lautoconus steindachneri (Hoernes, 1879)

Harzhauser, Mathias & Landau, Bernard, 2016, A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea, Zootaxa 4210 (1), pp. 1-178 : 93-95

Lautoconus steindachneri (Hoernes, 1879)

Figs 17 F, 21E1–E3, 21F1–F3

[ Dendroconus ] [ Conus ] Hochstetteri n. f.— Hoernes 1878a: 195 (nomen nudum) Conus (Dendroconus) Hochstetteri nov. form.— Hoernes & Auinger 1879: 24 (partim), pl. 3, fig. 3 [non Conus hochstetteri Martin, 1879 ].

Conus Steindachneri —Hoernes 1879: 201 [nov. nom. pro Conus hochstetteri Hoernes & Auinger, 1879 ]

Dendroconus Steindachneri nov. form.— Hoernes & Auinger 1879: pl. 3, plates captions.

Conus (Dendroconus) voeslauensis Hoernes und Auinger 1879 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 215, pl. 51, fig. 4 [non Kalloconus voeslauensis ( Hoernes & Auinger, 1879) ]

Conus (Cleobula) steindachneri Hoernes & Auinger, 1879 — Strausz 1966: 465, pl. 71, figs 6–9.

Conus (Dendroconus) steindachneri R. Hoernes & Auinger, 1879 — Mikuž 2009: 35, pl. 12, fig. 162.

Dendroconus steindachneri ( Hoernes et Auinger, 1879) — Kovács & Vicián 2013: 67, figs 47–49.

non Conus (Dendroconus) steindachneri Hoernes et Auinger—Csepreghy-Meznerics 1956: 422, pl. 12, figs 1–2.

non Conus (Dendroconus) steindachneri Hoernes et Auinger—Atanacković 1963: 79 , pl. 14, figs 5–6 [same specimen described as Conus steinabrunnensis in Atanacković, 1985 ]

non Conus (Cleobula) steindachneri Hörnes & Auinger, 1879 —Harzhauser 2002: 114, pl. 10, figs 1–3 [= Kalloconus ponderovagus (Sacco, 1893) ].

non Conus (Cleobula) steindachneri Hoernes et Auinger—Hasani & Vazari 2011: 128 , fig. 7/G.

Type material. Lectotype NHMW 1949 /0005/0004, Bad Vöslau ( Austria); illustrated in Hoernes & Auinger (1879, pl. 3, fig. 3) (designated herein); middle Miocene , Badenian (late Langhian).

Studied material. 1 spec. NHMW 1949 /0005/0004, Bad Vöslau ( Austria); illustrated in Hoernes & Auinger (1879, pl. 3, fig. 3); 15 spec . NHMW 1855 /0045/0858, Bad Vöslau ( Austria) ; 5 spec. NHMW 1997 z0178/1179, Bad Vöslau ( Austria) ; 3 spec. NHMW A1631, Lăpugiu de Sus ( Romania) ; 4 spec. NHMW 1856 /0004/0413, Lăpugiu de Sus ( Romania) .

Illustrated material. Figs 21 E1–E3: Bad Vöslau ( Austria), SL: 51.2 mm, MD: 32.1 mm, NHMW 1949/0005/ 0 0 0 4, illustrated in Hoernes & Auinger (1879, pl. 3, fig. 3); Figs 21 F1–F3: Bad Vöslau ( Austria), SL: 47.0 mm, MD: 30.5 mm, NHMW 1997z0178/1179; Fig. 17 F: Bad Vöslau ( Austria), SL: 41.0 mm, MD: 25.2 mm, NHMW 1855/ 0045/0858.

Revised description. Medium-sized, club-shaped solid shells with low conical spire. Pointed early spire; later spire whorls weakly convex, faintly striate; deep, narrowly canaliculated suture. Subsutural flexure shallow, weakly curved, strongly asymmetrical. Last spire whorl rapidly broadening; position of maximum diameter coinciding with periphery of prominent subangular to rounded shoulder; last whorl rapidly contracting, straightsided except for a very faint convexity mid-whorl. Prominent, strongly twisted siphonal fasciole; siphonal canal moderately wide, long, strongly recurved. Surface glossy with delicate growth lines and rather weak spiral grooves on lower third of last whorl. Colour pattern very weak (or weakly preserved) consisting of numerous thin spirals, which are continuous or composed of nearly continuous spirals of long dashes.

Shell ratios and measurement. n = 10 adult specimens: largest specimen: SL: 51.2 mm, MD: 33.4 mm, mean SL: 46.4 mm (σ = 3.7), mean MD: 29.4 mm (σ = 2.5), spire angle: µ = 116.9° (σ = 6.5°), last whorl angle: µ = 37.5° (σ = 1.0°), LW: µ = 1.58 (σ = 0.04), RD: µ = 0.72 (σ = 0.03), PMD: µ = 0.89 (σ = 0.03), RSH: µ = 0.12 (σ = 0.03).

Discussion. Hoernes & Auinger (1879) described this species as Conus hochstetteri in the text of their monograph. This name was already preoccupied by Conus hochstetteri Martin, 1879 , a Miocene species from Indonesia. The monograph of Martin was already available to R. Hoernes before publication of the monograph of Hoernes & Auinger (1879) but it was too late for changes in type setting. When recognizing the homonymy Hoernes (1879) introduced Conus steindachneri as new name for the Viennese species. This name was subsequently also added in the plate captions of Hoernes & Auinger (1879). Nevertheless, the authorship has to be passed to Hoernes (1879) and not to Hoernes & Auinger (1879) as frequently done in the literature.

When introducing this taxon Hoernes & Auinger (1879) referred to early Miocene specimens from Niederkreuzstetten ( Austria) and to middle Miocene specimens from Bad Vöslau ( Austria), thus mixing two species. The early Miocene specimens, including the specimen illustrated by Hörnes (1851, pl. 3, fig. 8), represent Kalloconus ponderovagus (Sacco, 1893) (see discussion of K. ponderovagus ). The description of Conus steindachneri by Hoernes & Auinger (1879) is mainly based on the middle Miocene specimens but the early Miocene specimens are syntypes as well. To settle this problem, we designate the specimen from Bad Vöslau, illustrated by Hoernes & Auinger (1879, pl. 3, fig. 3) as lectotype to clarify the status of this species.

Kalloconus moravicus ( Hoernes & Auinger, 1879) may develop a similar outline but differs in its slightly more curved subsutural flexure, the shorter siphonal canal and the wide-spaced spiral pattern. The specimen from the early Miocene of Iran, described by Hasani & Vazari (2011) as Conus steindachneri , is a different species with marked shoulder.

Paleoenvironment. In the Vienna and Transylvanian basins, this species occurs in offshore clays.

Distribution in Paratethys. Badenian (middle Miocene): Alpine-Carpathian Foredeep: Grund ( Austria); Vienna Basin: Bad Vöslau ( Austria), Mikulov-Kienberk ( Czech Republic) ( Hoernes & Auinger 1879); Pannonian Basin: Szob, Letkés ( Hungary) ( Strausz 1966; Kovács & Vicián 2013); Dacian Basin: Târnene, Orehovica ( Bulgaria) ( Kojumdgieva & Strachimirov 1960); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Hoernes & Auinger 1879; Boettger 1906); Krka Basin: Golobinjek pri Šentjerneju ( Slovenia) ( Mikuž 2009). The occurrence of this species in the Tortonian of Barcelona, mentioned by Faura I Sans (1908), needs confirmation.

Figures

FIGURE 21 A 1 – A 3. Lautoconus rotundus (Hoernes & Auinger, 1879), Steinebrunn (Austria), NHMW 1858 / 0015 / 0072, syntype. 21 B 1 – B 3. Lautoconus steinabrunnensis (Sacco, 1893), Grinzing (Austria), NHMW 1865 / 0001 / 0764. 21 C 1 – C 3. Lautoconus steinabrunnensis (Sacco, 1893), Steinebrunn (Austria), holotype, NHMW 1855 / 0045 / 0005. 21 D 1 – D 3. Lautoconus steinabrunnensis (Sacco, 1893), Lăpugiu de Sus (Romania), NHMW 1854 / 0035 / 0037. 21 E 1 – E 3. Lautoconus steindachneri (Hoernes, 1879), Bad Vöslau (Austria), NHMW 1949 / 0005 / 0004. 21 F 1 – F 3. Lautoconus steindachneri (Hoernes, 1879), Bad Vöslau (Austria), NHMW 1997 z 0178 / 1179. 21 G 1 – G 3. Lautoconus subraristriatus (Pereira da Costa, 1866), Enzesfeld (Austria), NHMW 1853 / 0010 / 0006 a. 21 H. Lautoconus subraristriatus (Pereira da Costa, 1866), Enzesfeld (Austria), NHMW A 1610.