Lautoconus miovoeslauensis (Sacco, 1893)

Harzhauser, Mathias & Landau, Bernard, 2016, A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea, Zootaxa 4210 (1), pp. 1-178 : 80-84

Lautoconus miovoeslauensis (Sacco, 1893)

Figs 17 B, 18D1–D3, 18E1–E3, 18F1–F2

Conus ventricosus Bronn—Hörnes 1851: 32 (partim), pl. 3, figs 6a–c [non Lautoconus ventricosus ( Gmelin, 1791) ].

Conus (Chelyconus) ventricosus Bronn—Hoernes & Auinger 1879: 49 , pl. 6, fig. 6 [non Lautoconus ventricosus ( Gmelin, 1791) ].

[ Conus (Chelyconus) tauroventricosus ] var. miovoeslauensis Sacc. — Sacco 1893b: 108 (nov. nom. pro Conus ventricosus in Hoernes & Auinger 1879 , pl. 6, fig. 6).

Conus (Chelyconus) ventricosus Bronn—Schaffer 1908: 98 , pl. 10, fig. 15 [non Lautoconus ventricosus ( Gmelin, 1791) ].

Conus ventricosus Brocc. —Friedberg 1911: 60, pl. 3, fig. 4 [non Lautoconus ventricosus ( Gmelin, 1791) ].

Chelyconus bitorosus Font. var. exventricosa Sacco—Schaffer 1912: 133 , pl. 49, fig. 4 (non figs 5–6).

Conus (Chelyconus) bitorosus exventricosus Sacco, 1893 — Hinculov 1968: 150, pl. 38, figs 1a–b [non Lautoconus exventricosus (Sacco, 1893) ].

Lautoconus bitorosus (Fontannes, 1880) — Landau et al. 2013: 239, pl. 38, figs 2–4, pl. 41, fig. 9, pl. 42, fig. 3, pl. 81, fig. 3.

non Conus (Chelyconus) miovoeslauensis Sacco, 1893 — Strausz 1966: 457, text-fig. 204.

non Conus ventricosus Bronn—Chira & Voia 2001: 156 , pl. 1, figs 1a–b, 4 a–b.

non Chelyconus miovoeslauensis (Sacco, 1893) — Kovács & Vicián 2013: 60, figs 11–15 [= Lautoconus pestensis nov. sp.]. non Monteiroconus bitorosus ( Fontannes, 1879) — Kovács & Vicián 2013: 76, figs 5–6, 82–86.

Type material. Holotype NHMW 1849 /0023/0003, Bad Vöslau ( Austria), illustrated in Hoernes & Auinger (1879, pl. 6, fig. 6); middle Miocene , Badenian (Langhian).

Studied material. Holotype and 10 spec. NHMW 1855 /0045/0857, Bad Vöslau ( Austria) ; 1 spec. NHMW 1937 /0002/0272, Bad Vöslau ( Austria), illustrated in Schaffer (1908, pl. 10, fig. 15); 4 spec . NHMW 2010 /0004/ 1459a-b Bad Vöslau ( Austria) ; 7 spec. NHMW 1846 /0037/0060, Steinebrunn ( Austria) ; 17 spec. NHMW 1846 / 0037/0062, Steinebrunn ( Austria) ; 7 spec. NHMW 1863 /0015/0398, Forchtenau ( Austria) ; 9 spec. NHMW 1870 / 0033/0003, Lăpugiu de Sus ( Romania) .

Illustrated material. Figs 18 D1–D3: holotype, Bad Vöslau ( Austria): SL: 51.3 mm, MD: 34.5 mm, NHMW 1849/0023/0003, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 6); Figs 18 E1–E3: Bad Vöslau ( Austria): SL: 65.7 mm, MD: 40.9 mm, NHMW 2010/0004/1459a; Figs 18 F1–F3: Bad Vöslau ( Austria): SL: 50.3 mm, MD: 30.8 mm, NHMW 2010/0004/1459b; Fig. 17 B: Bad Vöslau ( Austria): SL: 42.6 mm, MD: 25.6 mm, NHMW 1855/ 0045/0855.

Revised description. Medium-sized shells, spire coeloconoid, low to medium in height; very variable in height; spire whorls channelled, weakly striate to striate; more or less deep groove in mid-whorl. Subsutural flexure of medium depth, moderately curved, strongly asymmetrical; last whorl elongate and fig-shaped, rarely strongly ventricose. Shoulder high, rounded, sides convex, position of maximum diameter about ¼ of the way below the shoulder; very weakly constricted at base; bearing weak spiral grooves on the abapical third; aperture straight, widening and slightly flared abapically; siphonal canal short; siphonal fasciole weakly developed; colour pattern seen under UV light consisting of spirally arranged small irregular dashes, partly underlain by very thin spirals. Broad, wide-spaced flammulae on shoulder and last spire whorl.

Shell measurements and ratios. n = 17 adult specimens (only large morphs): largest specimen: SL: 65.7 mm, MD: 40.9 mm, mean SL: 51.2 mm (σ = 5.1), mean MD: 30.4 mm (σ = 3.6), spire angle: µ = 121.9° (σ = 8.6°), last whorl angle: µ = 38.7° (σ = 1.5°), LW: µ = 1.69 (σ = 0.08), RD: µ = 0.68 (σ = 0.02), PMD: µ = 0.86 (σ = 0.02), RSH: µ = 0.12 (σ = 0.03).

Discussion. Sacco (1893b) considered the Miocene specimen illustrated by Hörnes (1851, pl. 3, fig. 6) as Conus ventricosus to be conspecific with the Pliocene species Conus exventricosus Sacco, 1893 . He referred to the specimen as a typical representative, which could be read as designation as type specimen. Unfortunately, this specimen is lost. Later, Hoernes & Auinger (1879, pl. 6, fig. 6) illustrated another specimen from Bad Vöslau as Conus ventricosus . Their selection was somewhat unfortunate as Hoernes & Auinger (1879) illustrated an atypical low spired and broad specimen. Therefore, Sacco (1893b) introduced the varietal name miovoeslauensis for this specimen. The middle Miocene specimens from the Paratethys, like the Serravallian

specimens from the Turkish Karman Basin described by Landau et al. (2013), differ from the Pliocene Lautoconus bitorosus in their lower spires and more elongate shapes (see also Landau et al. 2013). Therefore, it might be justified to separate the Miocene specimens as Lautoconus miovoeslauensis (Sacco, 1893) .

This name was overlooked by Sieber (1958a) in his synthesis on Miocene gastropods of the Vienna Basin but was used by authors such as Strausz (1966) and Kovács & Vicián (2013). Unfortunately, the specimen illustrated by Strausz (1966, text-fig. 204) has a very broad last whorl and is not conspecific with L. miovoeslauensis . Similarly, the ovoid shells from Letkés in Hungary illustrated by Kovács & Vicián (2013) seem to represent a different species. The Italian specimen described by Sacco (1893b) as Conus exventricosus differs from the Paratethyan species in its rounded shoulder, the lower position of the maximum diameter and the more pronounced constriction of the base. Hall (1966) treated Conus miovoeslauensis as conspecific with Conus magnolapugyensis and both as synonyms of Lautoconus bitorosus (Fontannes, 1880) . Lautoconus miovoeslauensis differs from L. magnolapugyensis (Sacco, 1893) as redescribed herein in its smaller size, narrower and striate spire whorls, the distinct spiral threads on the base and the overall distinctly more slender outline.

Paleoenvironment. The pelitic sediment infill suggests offshore environments at least for specimens from Bad Vöslau; other localities, such as Steinebrunn, represent nearshore environments.

Distribution in Paratethys. Badenian (middle Miocene): Vienna Basin: Bad Vöslau, Baden, Steinebrunn ( Austria); Eisenstadt-Sopron Basin: Mattersburg, Forchtenau ( Austria); Alpine-Carpathian Foredeep: Grund ( Austria) ( Hoernes & Auinger 1879; Sieber 1949, 1956, 1958b), Korytnica ( Poland) (Friedberg 1911), Caransebeş-Mehadia Basin: Valea Bela Reca ( Romania) ( Hinculov 1968); Transylvanian Basin: Lăpugiu de Sus ( Romania) ( Hoernes & Auinger 1879).

The record of an early Miocene occurrence from Mörtersdorf (Austria) of Lower Austria mentioned by Schaffer (1910) and Mandic & Steiniger (2003) is based on poorly preserved specimens, which are almost certainly misidentified; note that the specimen illustrated by Schaffer (1912, pl. 49, fig. 4) derives from Bad Vöslau and is of middle Miocene age.

Proto-Mediterranean Sea and northeastern Atlantic. Serravallian (middle Miocene): Karaman Basin (Turkey) ( Landau et al. 2013). See Landau et al. (2013) for a detailed list of early Miocene to early Pleistocene occurrences of the closely related L. bitorosus (Fontannes, 1880) .

Figures

FIGURE 18 A 1 – A 3. Lautoconus magnolapugyensis (Sacco, 1893), Lăpugiu de Sus (Romania), NHMW 2016 / 0001 / 0002. 18 B 1 – B 3. Lautoconus magnolapugyensis (Sacco, 1893), Lăpugiu de Sus (Romania), NHMW 2016 / 0001 / 0001, holotype. 18 C 1 – C 2. Lautoconus magnolapugyensis (Sacco, 1893), Lăpugiu de Sus (Romania), NHMW 1870 / 0033 / 0002. 18 D 1 – D 3. Lautoconus miovoeslauensis (Sacco, 1893), Bad Vöslau (Austria), NHMW 1849 / 0023 / 0003, holotype. 18 E 1 – E 3. Lautoconus miovoeslauensis (Sacco, 1893), Bad Vöslau (Austria), NHMW 2010 / 0004 / 1459 a. 18 F 1 – F 3. Lautoconus miovoeslauensis (Sacco, 1893), Bad Vöslau (Austria), NHMW 2010 / 0004 / 1459 b.