Petrophilus Chaudoir, 1838

Sasakawa, Kôji & Kubota, Kôhei, 2006, Phylogenetic studies of the subgenus Petrophilus Chaudoir (Coleoptera: Carabidae: Pterostichus), with description of a new species sympatric with P. thunbergi Morawitz, Zootaxa 1357, pp. 31-43 : 35

Subgenus Petrophilus Chaudoir, 1838

Petrophilus Chaudoir, 1838: 9 , type species: Feronia findeli Dejean , by original designation. Jedliċka 1962: 260. Hůrka 1996: 278. Bousquet 2003: 505.

Euryperis Motschulsky, 1850 : ix, type species: Euryperis uralensis Motschulsky , by original designation. Jedliċka 1962: 255. Synonymy established by Kryzhanovskij et al. (1995).

Morphnosoma Lutshnik, 1915: 424 , type species: Carabus vulgaris Linné (= Carabus melanarius Illiger ), by original designation. Kryzhanovskij et al. 1995: 105. Hůrka 1996: 276. Bousquet 1999: 133. Arnett & Thomas 2001: 88. Bousquet 2003: 499. Sasakawa & Kubota 2005: 390. Syn. nov.

Euferonia Casey, 1918: 365 , type species: Feronia stygica Say , by original designation. Bousquet 1999: 135. Arnett & Thomas 2001: 88. Bousquet 2003: 494. Synonymy established, under the subgenus Morphnosoma , by Kryzhanovskij et al. (1995).

Feroperis Lafer, 1979: 5 , type species: Feronia jungens Tschitschérine , by original designation. Kryzhanovskij et al. 1995: 106. Bousquet 2003: 495. Synonymy established by Kryzhanovskij et al. (1995).

Moritapterus Berlov, 2000: 4, type species: Pterostichus thunbergi Morawitz , by original designation. Synonymy established, under the subgenus Morphnosoma , by Sasakawa & Kubota (2005).

Description. Body medium to large (minimum 10.0 mm: P. findeli [see Hùrka 1996]; maximum 20.0 mm: P. melanarius [see Bousquet 1999]). Head normal-sized. Pronotum wide and convex; laterobasal impressions single. Elytra oblong and convex; shoulders distinct. Male sternum 7 without sexual characteristics. Aedeagus stout, bent at approximately 90 degrees at the basal third. Endophallus large and strongly bent ventrally; gonopore weakly sclerotized, without a gonoporal piece; 3 lobes present in all species, one on the left ventrolateral surface near the gonopore, one on the left lateral surface near the gonopore, and one on the right ventrolateral surface near the gonopore; the surface of the lobe on the right ventrolateral surface near the gonopore sclerotized; the ventral surface with transverse pigmentation, except for P. l a c h r y m o s u s.