Bivetiella dertonensis ( Bellardi, 1841 )

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 18-19

Bivetiella dertonensis ( Bellardi, 1841)

Figs 3H, 5B 1–B 3, 5C 1–C 3, 5D 1–D 3

Cancellaria cancellata Lam. — Grateloup 1832: 339 partim [non Voluta cancellata Linnaeus, 1767 ].

C. cancellata Lam. — Grateloup 1846: pl. 25, fig. 10 [non Voluta cancellata Linnaeus, 1767 ].

Cancellaria Cancellata Lamarck var. Dertonensis Bellardi 1841: 28 , pl. 3, figs 13–14.

Cancellaria cancellata Lin. — Hauer 1837: 419 [non Voluta cancellata Linnaeus, 1767 ].

Cancellaria cancellata Lin. — Hörnes 1848: 19 [non Voluta cancellata Linnaeus, 1767 ].

Cancellaria subcancellata d’Orbigny 1852: 54 .

Cancellaria cancellata Lin. —Hörnes 1854: 316, pl. 34, figs. 20–22 [non Voluta cancellata Linnaeus, 1767 ].

Cancellaria cancellata Linn. — Hilber 1879: 30 [non Voluta cancellata Linnaeus, 1767 ].

Cancellaria subcancellata d’Orb. — Hoernes & Auinger 1890: 273.

Cancellaria cancellata Lam. — Rosiwal 1894: 83, 84 [non Voluta cancellata Linnaeus, 1767 ].

Cancellaria subcancellata d’Orb. — Boettger 1902: 38.

Cancellaria subcancellata d’Orb. — Boettger 1906: 49.

Cancellaria subcancellata d’Orb — Schaffer 1908: 110, pl. 11, fig 12.

Bivetia subcancellata d’Orb — Friedberg 1914: 244, pl. 15, fig. 7.

Cancellaria (Cancellaria) cancellata var. praecedens Beyrich — Sieber 1936: 76 [non Bivetiella praecedens ( Beyrich, 1856) ].

Cancellaria (Cancellaria) cancellata L. var. praecedens Beyr. — Sieber 1947: 157 [non Bivetiella praecedens ( Beyrich, 1856) ].

Cancellaria (Cancellaria) subcancellata perscalarata Sacco — Csepreghy-Meznerics 1954: 49, pl. 6, figs 19–21 [non Cancellaria (Bivetia) cancellata var. perscalarata Sacco, 1894 ].

Cancellaria (Cancellaria) cancellata praecedens Beyr. — Sieber 1956: 245 [non Bivetiella praecedens ( Beyrich, 1856) ].

Cancellaria (Cancellaria) subcancellata D’Orbigny — Csepreghy-Meznerics 1956: 415, pl. 9, figs 32–35

Cancellaria cancellata dertonensis Bellardi, 1841 — Strausz 1966: 376, pars, pl. 45, figs 5–6.

Cancellaria subcancellata perscalarata Sacco, 1894 — Strausz 1966: 376, pl. 44, figs 2–3 [non Cancellaria (Bivetia) cancellata var. perscalarata Sacco, 1894 ].

Cancellaria subcancellata infracosticillata (Sacco) — Csepreghy-Meznerics 1969b: 94, pl. 6, fig. 25 [non Cancellaria (Bivetia) cancellata var. infracostilicillata Sacco, 1894 ].

? Cancellaria (Cancellaria) cancellata praecedens Beyrich, 1856 — Steininger 1973: 433, pl. 7, fig. 12.

Cancellaria (Bivetiella) dertonensis Bellardi, 1841 — Davoli 1982: 21, pl. 1, figs 1–12.

Bivetia dertonensis ( Bellardi, 1841) — Ferrero Mortara et al. 1984: 169, pl. 32, fig. 4.

Cancellaria (Bivetiella) dertonensis Bellardi, 1841 — Davoli 1995: 226, pl. 1, figs 1–3.

Cancellaria (Bivetiella) neuvillei Peyrot, 1928 — Bałuk 1997: 42, pl. 14, fig. 7.

Cancellaria (Cancellaria) cancellata praecedens Beyrich — Schultz 1998: 70, pl. 28, fig. 12 [non Bivetiella praecedens ( Beyrich, 1856) ].

Bivetiella subcancellata B ( d’Orbigny, 1852)— Cahuzac et al. 2004: 227, pl. 8A–D.

non Cancellaria cancellata dertonensis Bellardi, 1841 — Strausz 1966: 376, pl. 45, figs 2–4 [= Contorta fenestrata ( Eichwald,

1830)].

Type material. syntype BS.045.09.010, Museo Regionale di Scienci Naturali , Torino ( Italy); Stazzano ( Italy), Tortonian (Late Miocene) [data from Ferrero-Mortara et al. (1984); Davoli (1982) discussed this specimen as potential lectotype without clear designation].

Studied material. 35 spec. Enzesfeld ( Austria) NHMW 1846 / 0037/0282, NHMW A1405 , NHMW A 2320 ; 25 spec. Bad Vöslau ( Austria) NHMW 2009 z0098/0018, NHMW 1874 /0024/0043; 23 spec. Gainfarn ( Austria) NHMW 1863 / 0015/1128, NHMW A384 ; 12 spec. Steinebrunn ( Austria) NHMW 1864 / 0001/0574; 1. spec. Pöls ( Austria) NHMW 1860 /0043/0001; 6 spec. Mikulov (Czech Rep.) NHMW 1851 /0010/0032; 4 spec. Lăpugiu de Sus ( Romania) NHMW 1875 /0009/0026.

Illustrated material. Figs. 3H, 5B 1–B 3: specimen illustrated in Hörnes (1854, pl. 34, figs 20a–b), Enzesfeld ( Austria), height: 32.4 mm, width: 20.4 mm, NHMW 1846/0037/0282b. Figs 5C 1–C 3: specimen illustrated in Hörnes (1854, pl. 34, figs 21a–b), Enzesfeld ( Austria), height: 28.8 mm, width: 19.6 mm, NHMW 1846/0037/ 0282a. Figs 5D 1–D 3: specimen illustrated in Hörnes (1854, pl. 34, figs 22a–b), Bad Vöslau ( Austria), height: 27.8 mm, width: 19.5 mm, NHMW 2009z0098/0018.

Discussion. This species was intensively discussed by Davoli (1982) and Landau et al. (2006; in prep.).

Distribution. A widespread species, which occured during the Middle Miocene from the North East Atlantic ( Peyrot 1928) via the Proto –Mediterranean Sea (Landau et al. in prep.) to the Paratethys, where it appears in all basins. It persisted into the Late Miocene in the North East Atlantic (Pereira da Costa 1867) and the Proto –Mediterranean Sea, where it is recorded up to the Messinian ( Davoli 1995). An occurrence from the Ottnangian (= middle Burdigalian) of Bavaria, described by Steininger (1973) as Cancellaria (Cancellaria) cancellata praecedens Beyrich, 1856 , is poorly preserved and its identity is unclear. It would be the earliest record of this species in the Paratethys.

Paratethys —Badenian: Polish–Carpathian Foredeep ( Poland: Korytnica); North Alpine Foreland Basin ( Austria: Grund, Windpassing), Vienna Basin ( Czech Republic: Mikulov: Austria: Steinebrunn, Gainfarn, Enzesfeld, Bad Vöslau, Baden, Baden–Sooss, Möllersdorf, Traiskirchen); Eisenstadt–Sopron Basin ( Austria: Forchtenau); Styrian Basin ( Austria: Pöls); Pannonian Basin ( Hungary: Szob, Letkés); Transylvanian Basin ( Romania: CoŞteiu de Sus). North East Atlantic —Serravallian ( France: Aquitaine Basin); Tortonian ( Portugal: Cacela Basin). Proto – Mediterranean Sea —Late Serravallian to Early Tortonian ( Turkey: Karaman Basin); Tortonian ( Italy: Po Basin); Messinian ( Italy: Po Basin).