Pseudobabylonella nysti (Hörnes, 1854) Harzhauser & Landau, 2012

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 13-15

Pseudobabylonella nysti (Hörnes, 1854) nov. comb.

Figs 3C, 4E 1–E 3

Cancellaria nysti Hörnes 1854: 305 , pl. 34, figs 1a–c.

Cancellaria (d. Narona ) nysti M. Hoern. — Hoernes & Auinger 1890: 280.

Admete nysti (M. Hö.) — Boettger 1906: 51.

Admete (Babylonella) fusiformis Cantr. var. nysti Hörn. — Meznerics 1932 /33: 346, pl. 13, figs 5a–b.

Admete (Babylonella) fusiformis Cantr. var. nysti Hörnes — Sieber 1936: 102.

Admete (Babylonella) fusiformis nysti Hörnes —Csepreghy–Meznerics 1956: 418, pl. 9, figs 28–29.

Cancellaria (Admete) fusiformis nysti Hörnes, 1856 [sic]— Strausz 1966: 384, pl. 12, figs. 13–14.

Type material. NHM Vienna ( Austria); see below for details .

Studied material. 8 spec. Steinebrunn ( Austria) NHMW 1887 / 0018/00044, NHMW 2009 z0098/0005; 3 spec. Grinzing ( Austria) NHMW 1864 /0001/0165; 2 spec. Möllersdorf ( Austria) NHMW 1862 /0001/0276; 22 spec. Szob ( Hungary) NHMW 1863 /0015/0311; 6 spec. Lăpugiu de Sus ( Romania) NHMW 1868 /0040/0184.

Illustrated material. Figs 3C, 4E 1–E 3: syntype illustrated in Hörnes (1854, pl. 34, figs 1a–c), Steinebrunn ( Austria), height: 9.6 mm, width: 3.9 mm, NHMW 2009 z0098/0005.

Discussion. Davoli (1982) and Janssen (1984a) highlighted the close relationship between this Paratethyan species and Cancellaria fusiformis Cantraine, 1835 , from the Miocene to Pliocene of the North Sea Basin, the North East Atlantic and the Mediterranean Sea. This species was placed into Pseudobabylonella by Brunetti et al. (2009), which also requires including the Paratethyan shell within that genus. Pseudobabylonella fusiformis differs constantly in its broader shell, the usually more convex whorls and the coarser sculpture. A further difference is the presence of lirae in the inner side of the outer lip of P. nysti , whereas the lip of P. fusiformis is smooth (cf. Landau et al. 2006). Finally, the protoconch of P. nysti is much higher and its sutures are less incised (cf. Fig. 4a in Landau et al. 2006).

Distribution. Known so far only from the early and middle Badenian of the Paratethys Sea. Paratethys —Badenian: Vienna Basin ( Czech Republic: Lissitz; Austria: Steinebrunn, Grinzing, Möllersdorf,

Perchtoldsdorf); Mecsek Mountains and Pannonian Basin ( Hungary: Szob); Transylvanian Basin ( Romania:

Lăpugiu de Sus, CoŞteiu de Sus).