Sveltia dertovaricosa ( Sacco, 1894 )

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 49-50

Sveltia dertovaricosa ( Sacco, 1894)

Figs 3S, 9H 1–H 3

Cancellaria varicosa Brocc. — Hauer 1837: 419 [non Voluta varricosa Brocchi, 1814 ].

Cancellaria varicosa Brocc. — Hörnes 1848: 19 [non Voluta varricosa Brocchi, 1814 ].

Cancell. varicosa Brocchi — Eichwald 1853: 200 [non Voluta varricosa Brocchi, 1814 ].

Cancellaria varicosa Brocc. —Hörnes 1854: 309, pl. 34, figs 6a–b [non Voluta varricosa Brocchi, 1814 ].

Cancellaria (a. Narona ) varicosa Brocc. — Hoernes & Auinger 1890: 278 [non Voluta varricosa Brocchi, 1814 ].

Cancellaria (Sveltia) dertovaricosa Sacco, 1894: 57 , pl. 3, fig. 49.

Cancellaria (Sveltia) varicosa var. miocenica Sacco, 1894: 56 , pl. 3, fig. 47.

Cancellaria (Sveltia) varicosa Brocc. – Boettger 1902: 39 [non Voluta varricosa Brocchi, 1814 ].

Cancellaria (Sveltia) varicosa Brocc. – Boettger 1906: 51 [non Voluta varricosa Brocchi, 1814 ].

Sveltia dertovaricosa Sacco — Friedberg 1914: 247, pl. 15, fig. 10.

Sveltia dertovaricosa var. subasuturata Sacco — Friedberg 1914: 247, pl. 15, fig. 11.

Sveltia varicosa (Brocchi) var. simplicior Sacco — Peyrot 1928: 217, pl. 14, figs 5–6.

Sveltia varicosa (Broc.) mut. bearnensis nov. mut.— Peyrot 1928: 219, pl. 13, fig. 8.

Sveltia varicosa Brocc. — Janoschek 1932: 76 [non Voluta varricosa Brocchi, 1814 ].

Cancellaria (Sveltia) dertovaricosa Sacco — Sieber 1936: 93, pl. 3, figs 1a–b, 8a–b.

Cancellaria (Sveltia) varicosa Brocchi var. miocenica Sacco — Sieber 1936: 94.

Cancellaria (Sveltia) dertovaricosa Sacco — Sieber 1947: 157.

Cancellaria dertovaricosa Sacco — Csepreghy-Meznerics 1950: 59, pl. 3, fig. 14.

Cancellaria uniangulata Deshayes — Csepreghy-Meznerics 1950: 58, pl. 3, fig. 17 [non Cancellaria uniangulata Deshayes, 1830 ].

Sveltia (Sveltia) dertovaricosa Sacco — Erünal-Erentöz 1958: 87, pl. 13, figs 11–12.

Sveltia (Sveltia) dertovaricosa Sacco, 1890 [sic]— Kojumdgieva & Strachimirov 1960: 163, pl. 42, fig. 15.

Cancellaria (Sveltia) varicosa dertovaricosa Sacco, 1894 — Strausz 1966: 380, pl. 43, figs 10–12.

? Narona (Sveltia) dertovaricosa (acco) [sic]— Hölzl 1973: 194.

? Narona (Sveltia) varicosa miocenica (Sacco) — Hölzl 1973: 194.

Narona (Sveltia) dertovaricosa ( Sacco, 1894) — Davoli 1982: 39, pl. 4, figs 1–7, 10–12.

Narona (Sveltia) varicosa miocaenica [sic] ( Sacco, 1894)— Švagrovský 1982: 401, pl. 4, fig. 6.

Narona (Sveltia) varicosa ( Brocchi, 1814) — Davoli 1995: 243 (partim), pl. 5, figs 1–5, 18, pl. 6, fig. 2 [Miocene specimens only].

Narona (Sveltia) varricosa ( Brocchi, 1814) — Bałuk 1997: 48, pl. 16, figs 6–8 [non Voluta varricosa Brocchi, 1814 ].

Narona (Sveltia) varicosa miocenica Sacco — Schultz 1998: 72, pl. 29, figs 4a–b.

Narona (Sveltia) varricosa ( Brocchi, 1814) — Mikuž 2009: 28, pl. 9, figs 118–120 [non Voluta varricosa Brocchi, 1814 ].

Sveltia dertovaricosa Sacco, 1894 — Brunetti et al. 2011: 94, fig. 4G–H, fig. 5A–B.

Type material. syntype BS.045.13.010, Museo Regionale di Scienci Naturali , Torino ( Italy); Stazzano ( Italy), Tortonian (Late Miocene) [data from Ferrero-Mortara et al. 1984] .

Studied material. 20 Grund ( Austria) NHMW A891; 19 spec. Kalladorf ( Austria) NHMW A389 ; 17 Steinebrunn ( Austria) NHMW A1410 ; 10 Niederleis ( Austria) NHMW 1865 / 0001/0623; 20 Enzesfeld-Gainfarn-Steinebrunn (mixed lot) ( Austria) NHMW 1846 /0037/0292; 5 spec. Enzesfeld ( Austria) NHMW 446 ; 21 spec. Bad Vöslau ( Austria) NHMW 1872 /0030/0104; 32 spec. Lăpugiu de Sus ( Romania) NHMW 1870 /0033/0032; 21 spec. Bujtur ( Romania) NHMW 1862 /0001/0547; 29 spec. Korytnica ( Poland) NHMW 1853 /0038/0029.

Illustrated material. Fig: 3S: Enzesfeld ( Austria) height: 25.6 mm, width: 12.4 mm NHMW 446. Figs 9H 1– H 3: specimen illustrated in Hörnes (1854, pl. 34, figs 6a–b), Enzesfeld ( Austria), height: 39.7 mm, width: 19.3 mm, NHMW 1846/0037/0292.

Discussion. The specimen illustrated by Hörnes (1854) as Cancellaria varicosa Brocc. is extraordinarily large and angulated. Most other shells range between 20–30 mm in height and have convex whorls with numerous, varix-like axial ribs, crossed by a dense spiral sculpture, comprising a quite regular succession of primary cords with an intercalated secondary thread. As pointed out by Landau et al. (2006), the distinctly less angulated and smaller Miocene shells, which develop a weak third columellar fold and lack the inflated last whorl of Sveltia varicosa , have to be treated as S. dertovaricosa ( Sacco, 1894) [see Landau et al. 2006 for a discussion on the use of the species name varricosa versus varicosa ].

Distribution. The earliest representatives of this species appear during the Early Miocene of the Proto –Mediterranean Sea, where it persists up to the Late Miocene. During the Middle Miocene it spreads also into the Paratethys and the North–East Atlantic. It might be present in the Paratethys already during the Early Miocene as indicated by Hölzl (1973) for the Ottnangian of Bavaria in a list without descriptions.

North East Atlantic —Langhian and Serravallian: Aquitaine Basin ( France). Proto –Mediterranean Sea —Burdigalian: Colli Torinesi ( Italy: no locality given in Sacco, 1894); Late Serravallian to Early Tortonian: Karaman Basin ( Turkey); Tortonian: Po Basin ( Italy). Paratethys —Badenian: Polish–Carpathian Foredeep ( Poland: Korytnica, Zglobice); North Alpine Foreland Basin ( Austria: Grund, Kalladorf, Windpassing); Vienna Basin ( Austria: Enzesfeld, Gainfarn, Steinebrunn, Bad Vöslau, Baden); Styrian Basin ( Austria: Ritzing); Transylvanian Basin ( Romania: Lăpugiu de Sus, CoŞteiu de Sus, Bujtur); Krka Basin ( Slovenia: Gorenje Mokro Polje); Dacian Basin ( Bulgaria: Opanec).