Sveltia lyrata ( Brocchi, 1814 )

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 51-52

Sveltia lyrata ( Brocchi, 1814)

Figs 3T–U, 10A 1–A 3, 10B 1–B 2

Voluta lyrata Brocchi, 1814: 311 , pl. 3, fig. 6.

Cancellaria lyrata Bronn —Pusch 1837: 130.

Cancellaria lyrata Brocc. — Hörnes 1848: 19.

Cancellaria lyrata Brocc. —Hörnes 1854: p. 308, pl. 34, figs 4–5.

Cancellaria (q. Trigonostoma ) lyrata Brocc. — Hoernes & Auinger 1890: 278.

Cancellaria (Sveltia) lyrata var. badensis Sacco, 1894: 60 .

Cancellaria (Sveltia) lyrata var. vindobonensis Sacco, 1894: 62 .

Cancellaria (Trigonostoma) lyrata Brocc. – Boettger 1902: 39.

Cancellaria (Trigonostoma) lyrata Brocc. – Boettger 1906: 51.

Cancellaria subcancellata d’Orb — Schaffer 1908: 109, pl. 11, fig. 11.

Cancellaria (Sveltia) lyrata Brocchi – Sieber 1936: 96.

Narona (Sveltia) lyrata (Brocc.) — Sieber 1936: 245.

Cancellaria (Sveltia) lyrata Brocchi —Czepreghy-Meznerics 1956: 416, pl. 9, figs 22–23.

Sveltia (Calcarata) lyrata ( Brocchi 1814) — Kojumdgieva & Strachimirov 1960: 162, pl. 42, fig. 18.

Cancellaria (Calcarata) lyrata Brocchi 1814 — Strausz 1966: 382, pl. 43, figs 19–24.

Cancellaria (Calcarata) lyrata spinulosa Brocchi 1814 — Strausz 1966: 382, pl. 45, fig. 1.

Cancellaria (Narona) lyrata Brocchi — Csepreghy-Meznerics 1972: 31, pl. 14, figs 21, 24.

Cancellaria (Narona) lyrata spinulosa (Brocchi) — Csepreghy-Meznerics 1972: 31, pl. 14, figs 25, 27, 29.

Sveltia lyrata ( Brocchi, 1814) — Landau et al. 2006: 75, pl. 6, fig. 1 [cum syn].

Sveltia lyrata ( Brocchi, 1814) — Brunetti et al. 2011: 96, fig. 5E–H, fig. 6A–F.

non Sveltia lyrata Brocc. — Friedberg 1914: 246, pl. 15, fig. 9 [= Calcarata vindobonensis ( Hilber, 1892) ].

non Narona (Sveltia) lyrata ( Brocchi, 1814) — Atanacković 1985: 165, pl. 37, figs 3–4 [= Calcarata vindobonensis ( Hilber, 1892) ].

Type material. syntype, Inv. Nr. 89, Museo Civico di Storia Naturale di Milano ( Italy); Parlascio ( Italy), Pliocene [data from Rossi-Ronchetti 1955 and Verhecken 2007] .

Studied material. 3 spec. Grund ( Austria) NHMW 2009 z0098/0007, NHMW 1851 /0002/0050; 26 spec. Baden ( Austria) NHMW 1846 / 0037/0290; 35 Baden–Sooss ( Austria) NHMW 1866 / 0001/0649; NHMW 1872 / 0030/0058; NHMW A1412 ; 3 spec. Traiskirchen ( Austria) NHMW A2223 ; 5 spec. Hrušovany (Czech Rep.) NHMW 1863 /0015/0968; 29 spec. Lăpugiu de Sus ( Romania) NHMW 1854 / 0030/0230, NHMW 1865 /0001/ 0212.

Illustrated material. Fig. 3T: Baden ( Austria), height: 24.3 mm, width: 14.9 mm, NHMW 1846 /0037/0290. Fig : 3U: Baden ( Austria), height: 18.5 mm, width: 11.6 mm, NHMW 1846 /0037/0290. Figs 10A 1–A 3: specimen illustrated in Hörnes (1854, pl. 34, figs 4a–b), Baden ( Austria), height: 32.7 mm, width: 19.4 mm, NHMW 1846 / 0037/0290, holotype of Cancellaria badensis Sacco, 1894 .

Figs 10B 1–B 2: specimen illustrated in Hörnes (1854, pl. 34, figs 5a–b), Grund ( Austria), height: 32.5 mm, width: 18.2 mm, NHMW 2009z0098/0007, holotype of Cancellaria vindobonensis Sacco, 1894 [non Cancellaria vindobonensis Handmann, 1892 ].

Discussion. Sveltia lyrata ( Brocchi, 1814) is quite a common species in deeper water settings, displaying a wide variability in sculpture. Especially the spiral sculpture ranges from prominent to zero, resulting in the presence or absence of a weak row of spines close below the shoulder. Even the umbilicus ranges from nearly absent to distinct, bordered by a fasciole. A comparable variability is documented by Chirli (2002) for Pliocene specimens from Italy. Therefore, a specific separation of the Middle Miocene from the Pliocene shells seems to be unjustified. Consequently, the variety names badensis and vindobonensis , proposed by Sacco (1894), are considered synonyms. Moreover, Cancellaria vindobonensis Sacco, 1894 is a junior primary homonym of Cancellaria vindobonensis Hilber, 1892 .

This species is supposed to be still present along the coast of western Africa ( Verhecken 2007). The extant shells are more slender and develop a conspicuous spiral sculpture with primary and secondary ribs covering the entire shell. Another typical feature is the spinate angulation on the aperture. In our opinion, the extant representatives could belong to a tropical Atlantic offshoot of Sveltia lyrata .

Distribution. Landau et al. (2006) gave an extensive summary of the distribution of that species with numerous references. According to their synopsis, this widespread species appears already during the Early Miocene in the North Sea Basin and the Proto –Mediterranean Sea and becomes ubiquitous in the Middle Miocene, documented from the North Sea, the North East Atlantic, the Proto –Mediterranean and the Paratethys. Afterwards it seems to be limited to the Proto –Mediterranean, where it finally disappears during the Middle Pliocene.

Paratethys —?Ottnangian: North Alpine Foreland Basin (Bavaria: Kaltenbachgraben); Badenian: North Alpine Foreland Basin ( Austria: Grund); Vienna Basin ( Czech Republic: Hrušovany; Austria: Gainfarn, Grinzing, Bad Vöslau, Baden–Sooss, Baden, Möllersdorf, Traiskirchen); Eisenstadt–Sopron Basin ( Austria: Forchtenau; Hungary: Sopron); Bükk and Mecsek Mountains and Pannonian Basin ( Hungary: Devecser, Borsod, Szob, Balaton); Transylvanian Basin ( Romania: Lăpugiu de Sus, CoŞteiu de Sus); Dacian Basin ( Bulgaria: Staropatica). North Sea —Early Miocene ( Belgium: Antwerp); Middle Miocene ( Germany, Denmark, Netherlands). North East Atlantic —Middle Miocene ( France: Aquitaine Basin). Proto –Mediterranean Sea —Early Miocene ( Italy: Colli Torinesi, Baldissero, Sciolze); Late Miocene ( Italy: Montegibbio; Spain); Early Pliocene ( Italy, Spain, France); Middle Pliocene ( Italy); maybe also in the Pliocene Mediterranean Sea and Recent (western Africa).