Scalptia spinosa ( Grateloup, 1827 )

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 41-42

Scalptia spinosa ( Grateloup, 1827)

Figs 3O, 8F 1–F 3, 8G 1–G 3, 8H 1–H 3

Cancellaria Spinosa Grateloup 1827: 21 .

Cancellaria spinifera Grateloup 1832: 342 .

Cancellaria spinifera Grateloup 1846 : pl. 25, fig. 15.

Cancellaria spinifera Grat. —Hörnes 1854: 323, pl. 35, figs. 6–8.

Cancellaria (h. Trigonostoma ) spinifera Grat. — Hoernes & Auinger 1890: 277.

Cancellaria (Gulia) geslini var. crassopostica Sacco 1894: 23 , pl. 2, fig. 10.

Cancellaria (Gulia) geslini var. compressohirta Sacco 1894: 23 , pl. 2, fig. 11.

Cancellaria (Gulia) geslini var. hirtissima Sacco 1894: 24 .

Cancellaria (Trigonostoma) spinifera Grat. – Boettger 1902: 39.

Cancellaria (Gulia) geslini (Bast.) var. compressohirta Sacco — Boettger 1906: 50.

Trigonostoma spiniferum Grateloup — Peyrot 1928: 243, pl. 13, figs 32–33.

Trigonostoma spinifera Grat. — Janoschek 1932: 76.

Cancellaria (Trigonostoma) spinifera Grat. — Sieber 1936: 84, pl. 3, fig. 10.

Cancellaria (Trigonostoma) spinifera Grat. — Sieber 1947: 157.

Trigonostoma (Trigonostoma) spiniferum (Grat.) — Sieber 1956: 245.

Cancellaria (Trigonostoma) spinifera Grateloup — Strausz 1966: 377 (partly).

Cancellaria (Trigonostoma) spinifera Grat. — Csepreghy-Meznerics 1972: 31, pl. 14, fig. 28.

Trigonostoma (Trigonostoma) spiniferum (Grateloup, 1845) — Davoli 1982: 33, pl. 1, figs 10, 13–14, 20.

Gulia geslini compressohirta Sacco — Ferrero Mortara et al. 1984: 164, pl. 30, fig. 9.

Trigonostoma spiniferum (Grateloup, 1845) — Bałuk 1997: 46, pl. 15, fig. 1.

Trigonostoma (Trigonostoma) spiniferum (Grateloup) — Schultz 1998: 72, pl. 29, figs 2–3b.

Scalptia (s.l.) spinosa (Grateloup, 1927) — Cahuzac et al. 2004: 234, figs 3A–K.

Scalptia spinosa (Grateloup, 1927) — Brunetti et al. 2011: 126, fig. 23A–G.

non Trigonostoma cf. Michelini Bell.— Friedberg 1914: 250, text–fig. 55 [= Scalptia exgeslini ( Sacco, 1894) ]. non Cancellaria (Trigonostoma) spinifera Grat. — Csepreghy-Meznerics 1950: 58, pl. 3, fig. 13 [= Scalptia exgeslini ( Sacco,

1894)]. non Cancellaria (Trigonostoma) spinifera Grateloup — Hölzl 1958: 255, pl. 21, fig. 10 [= Gulia geslini ( Basterot, 1825) ]. non Cancellaria (Trigonostoma) spinifera Grateloup (1832) 1845 — Strausz 1966: 377, pl. 45, fig. 14 [= Scalptia exgeslini

( Sacco, 1894)].

Type material. lectotype 65-2-74 designated by Cahuzac et al. (2004), Université de Bordeaux-1 ( France); Saint- Jean-de-Marsacq ou Saubrigues ( France), Burdigalian or Langhian (Early or Middle Miocene) .

Studied material. 9 Grund ( Austria) NHMW A895 ; 11 spec. Baden ( Austria) NHMW 1847 / 0037/0075; 47 spec. Steinebrunn ( Austria) NHMW 1846 /0037/0288, NHMW 1867 /0002/3937, NHMW A1401 , NHMW A381 ; 10 Enzesfeld-Gainfarn (mixed lot) ( Austria) NHMW 1846 /0037/0258; 16 spec. Mikulov (Czech Rep.) NHMW 2009 z0098/0019, NHMW 1849 /0012/0012, NHMW A2321 ; 41 spec. Lăpugiu de Sus ( Romania) NHMW 1868 / 0001/0469, NHMW A1403 .

Illustrated material. Fig. 3O: Baden ( Austria), height: 23.1 mm, width: 16.1 mm, NHMW 1847 / 0037/0075. Figs 8H 1–H 3: specimen illustrated in Hörnes (1854, pl. 35, figs 6a–c), Mikulov ( Czech Republic), height: 38.5 mm, width: 25.8 mm, NHMW 2009 z0098/0019. Figs 8G 1–G 3: specimen illustrated in Hörnes (1854, pl. 35, figs 8a–c), Baden ( Austria), height: 22.2 mm, width: 16.5 mm, NHMW 1847 / 0037/0075, syntype of Cancellaria hirtissima Sacco, 1894 . Figs 8F 1–F 3: specimen illustrated in Hörnes (1854, pl. 35, figs 7a–c), Steinebrunn ( Austria), height: 29.2 mm, width: 22.3 mm, NHMW 1846 /0037/0288.

Discussion. A very widepread species, which is quite variable concerning its spiny sculpture. Specimens from deeper marine settings tend to be more depressed and develop prominent, partly open spines along the shoulder. Comparable morphs, however, appear also in shallow water assemblages, where the robust morphs without spines predominate. Thus, a separation of the spiny morph as hirtissima , as proposed by Sacco (1894), is unjustified. The specimen illustrated by Hörnes (1854, pl. 35) as fig. 6 is an extreme morph with slightly worn surface and poorly developed shoulder. The well-preserved protoconch figured ( Fig. 3O) clearly illustrates a dome-shaped multispiral protoconch with a small nucleus.

This species was usually referred to as Cancellaria spinifera Grateloup, 1832 . Cahuzac et al. (2004), however, documented that C. spinosa Grateloup 1827 has priority.

Distribution. A widespread species, which appears throughout the Miocene in the North East Atlantic and the Proto –Mediterranean ( Bellardi 1841; Cahuzac et al. 2004; Pereira da Costa 1867; Davoli 1982; Landau et al. in prep.). During the Middle Miocene it spreads also into the Paratethys. Its last occurrence is known from the Early Pliocene of Italy ( D’Ancona 1872).

North East Atlantic —Burdigalian and Langhian (Aquitaine Basin, France); Late Miocene (Cacela Basin, Portugal). Paratethys —Badenian: Polish–Carpathian Foredeep ( Poland: Korytnica); North Alpine Foreland Basin ( Austria: Grund, Windpassing), Vienna Basin ( Czech Republic: Mikulov; Austria: Steinebrunn, Gainfarn, Enzesfeld, Bad Vöslau, Baden–Sooss, Baden); Eisenstadt–Sopron Basin ( Austria: Forchtenau); Styrian Basin ( Austria: Ritzing); Mecsek Mountains ( Hungary); Transylvanian Basin ( Romania: Lăpugiu de Sus, CoŞteiu de Sus). Proto – Mediterranean Sea –Burdigalian (Colli Torinesi, Italy); Late Serravallian–Early Tortonian (Karaman Basin, Turkey); Tortonian (Po Basin, Italy); Early Pliocene: (Central Mediterranean, Italy).