Scalptia dertocosticillata ( Sacco, 1894 ) Harzhauser & Landau, 2012

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 37-38

Scalptia dertocosticillata ( Sacco, 1894) nov. comb.

Cancellaria (Trigonostoma) imbricatum var. dertocosticillata Sacco, 1894: 13 ; pl. 1, fig. 31.

Trigonostoma puschi R. Hoern. i Auing.— Friedberg 1914: 248, pl. 15, fig. 12.

Trigonostoma puschi (R. Hoernes et Auinger, 1879) — Zelinskaya et al. 1968: 216, pl. 49, figs 33–34.

Trigonostoma imbricatum var. dertocosticillata Sacco, 1894 — Ferrero Mortara et al. 1984: 161, pl. 29, fig. 3.

Trigonostoma puschi ( Hoernes & Auinger, 1890) — Bałuk 1997: 45, pl. 15, figs 4–5.

Type material. syntype BS.045.01.030, Museo Regionale di Scienci Naturali , Torino ( Italy); Stazzano ( Italy), Tortonian (Late Miocene) [data from Ferrero-Mortara et al. 1984] .

Discussion. This species was frequently intermingled with Scalptia polonica (Pusch, 1837) . The specimens presented by Bałuk (1997), however, are more similar to Scalptia dertocosticillata ( Sacco, 1894) from the Tortonian of Italy. Both species develop a comparable outline with a slight concavity below the shoulder and a somewhat inflated lower third of the last whorl. A marked shoulder angulation causes a terraced spire with deep cavities between the axial ribs on the narrow shelf. The axial ribs are prominent on the spire whorls and become slightly less distinct on the last whorl. The spiral cords cross the axials, and a secondary spiral rib is intercalated in the interspaces, giving the surface a slightly scaley appearance.

Sacco (1894) described this species as variety of Cancellaria imbricata Hörnes, 1854 , which develops a much wider umbilicus. Scalptia burdigalensis ( Peyrot, 1928) , from the Burdigalian of the Aquitaine Basin, might be an ancestor of the Paratethyan species. It differs mainly in the wider umbilicus and lacks the secondary spiral ribs (see Cahuzac et al. 2004). The placement of this species in the genus Scalptia is based mainly on the similarity with this French species but will need further confirmation.

Despite the morphologic similarity between the Paratethyan species and the Italian type, doubts remain if they are really conspecific. The Paratethyan shells are known so far only from the Polish–Carpathian Foredeep, which was repeatedly isolated from other Paratethyan basins, resulting in an increased endemism (e.g. within nassariids Harzhauser & Kowalke 2004). Thus, the similarity with Scalptia dertocosticillata ( Sacco, 1894) might represent a convergence as well.

A dubious Early Miocene record is mentioned by Steininger et al. (1971) from Lipovany in Slovakia as Cancellaria (Trigonostoma) puschi . The authors refer to the specimens illustrated by Friedberg (1914), which are considered herein as Scalptia cf. dertocosticillata ( Sacco, 1894) . The whereabouts of the Slovak material are unclear and the record lacks a description and illustration. Therefore, we refrain from listing it in the chresonynmy.

Distribution. Known from the Badenian of the north–eastern Paratethys and from the Middle–Late Miocene of the Proto –Mediterranean Sea.

Paratethys —Badenian: Polish–Carpathian Foredeep ( Poland: Korytnica, Piṅczów, Zglobice). Proto –Mediterranean Sea —Tortonian: Po Basin ( Italy: Stazzano, Montegibbio, Sant’Agata). Late Serrvallian–Early Tortonian: Karaman Basin ( Turkey, BL coll.).